Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rhombophryne maraorao Scherz, Glaw, Petzold, Andreone, Köhler, Raselimanana, Hutter, Belluardo, Rakotoarison, Hofreiter, Vences, and Crottini, 2026

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Cophylinae > Genus: Rhombophryne > Species: Rhombophryne maraorao

Rhombophryne maraorao Scherz, Glaw, Petzold, Andreone, Köhler, Raselimanana, Hutter, Belluardo, Rakotoarison, Hofreiter, Vences, and Crottini, 2026, Vert. Zool., Senckenberg, 76: 520. Holotype: MRSN A2115, by original designation. Type locality: "Ambatoledama Corridor (Andasin’i Governera) (ca 15.30°S, 50.02°E, 650 m a.s.l.), Masoala Peninsula, Sava Region, northeastern Madagascar". ZooBank Publication registration: 72D3AFA4-D4DA-4EF8-9734-B592172E6095

Common Names

None noted.

Distribution

Known from (1) Ambatoledama Corridor (Andasin’i Governera); (2) Amparihy, (3) Ambolokopatrika-Betaolana, and possibly (4) Anandrivola. Referral of material from Marojejy to this species remains too uncertain to include in its distribution at present. Elevation ca 650–950 m.

Geographic Occurrence

Natural Resident: Madagascar

Endemic: Madagascar

Comment

In the Rhombophryne testudo group of Scherz, Glaw, Petzold, Andreone, Köhler, Raselimanana, Hutter, Belluardo, Rakotoarison, Hofreiter, Vences, and Crottini, 2026, Vert. Zool., Senckenberg, 76: 485–610, who provided a species account covering phylogenetic relationships, morphology, osteology, natural history, and other relevant literature.

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