Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Uperoleia rugosa (Andersson, 1916)

Class: Amphibia > Order: Anura > Superfamily: Myobatrachoidea > Family: Myobatrachidae > Genus: Uperoleia > Species: Uperoleia rugosa

Pseudophryne rugosa Andersson, 1916, K. Svensk. Vetensk. Akad. Handl., 52: 13. Holotype: NHRM 1630, according to Parker, 1940, Novit. Zool., 42: 71, Cogger, Cameron, and Cogger, 1983, Zool. Cat. Aust., Amph. Rept., 1: 34, and Tyler, 1985, in Frost (ed.), Amph. Species World: 408. Type locality: "[Mt.] Colosseum, Southern Queensland", Australia.

Pseudophryne fimbrianus Parker, 1926, Ann. Mag. Nat. Hist., Ser. 9, 17: 669. Holotype: BMNH 1923.11.12.3, according to Parker, 1940, Novit. Zool., 42: 71; renumbered as BMNH 1947.2.18.70 according to Cogger, Cameron, and Cogger, 1983, Zool. Cat. Aust., Amph. Rept., 1: 32, and Tyler, 1985, in Frost (ed.), Amph. Species World: 407. Type locality: "St. George district, Queensland (Lat. 28° 14′ S., Long. 148° 35′ E.)", Australia. Synonymy by Loveridge, 1935, Bull. Mus. Comp. Zool., 78: 7 (with Uperoleia marmorata rugosa); and by Davies and Littlejohn, 1986, Trans. R. Soc. S. Aust., 110: 113.

Uperoleia marmorata rugosa — Loveridge, 1935, Bull. Mus. Comp. Zool., 78: 7, 31.

Uperoleia fimbrianus — Tyler, Davies, and Martin, 1981, Aust. J. Zool., Suppl. Ser., 29 (79): 18.

Uperoleia rugosa — Tyler, Davies, and Martin, 1981, Aust. J. Zool., Suppl. Ser., 29 (79): 18.

Prohartia fimbrianus — Wells and Wellington, 1985, Aust. J. Herpetol., Suppl. Ser., 1: 3. Disagreement in gender.

Prohartia rugosa — Wells and Wellington, 1985, Aust. J. Herpetol., Suppl. Ser., 1: 3. Elevation to species status without discussion.

Uperoleia capitulata Davies, McDonald, and Corben, 1986, Proc. R. Soc. Victoria, 98: 163. Holotype: QM J26428, by original designation. Type locality: "Bollon, Queensland (28°02′ [S], 147°29′ [E])", Australia. Synonymy by Catullo and Keogh, 2014, Mol. Phylogenet. Evol., 79: 115.

English Names

Red-groined Toadlet (Cogger, 1975, Rept. Amph. Australia: 83),

Chubby Gungan (Ingram, Nattrass, and Czechura, 1993, Mem. Queensland Mus., 33: 223).

Mud Toadlet (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 96).

Wrinkled Toadlet (Barker, Grigg, and Tyler, 1995, Field Guide Aust. Frogs., Ed. 2: 339; Tyler and Knight, 2009, Field Guide Frogs Aust.: 148).

Desert Gungan (Uperoleia capitulata [no longer recognized]: Ingram, Nattrass, and Czechura, 1993, Mem. Queensland Mus., 33: 223).

Grass Toadlet (Uperoleia capitulata [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 95).

Small-headed Toadlet (Uperoleia capitulata [no longer recognized]: Barker, Grigg, and Tyler, 1995, Field Guide Aust. Frogs., Ed. 2: 320; Tyler and Knight, 2009, Field Guide Frogs Aust.: 148).

Distribution

Northeastern South Australia to southern Queensland and to including all but the southern part of eastern New South Wales, Australia.

Comment

Tyler, Davies, and Martin, 1981, Aust. J. Zool., Suppl. Ser., 29 (79): 17, redefined the species and questioned the identity of literature records. Subsequently, Davies and Littlejohn, 1986, Trans. R. Soc. S. Aust., 110: 111–143, redelimited the species. Robertson, 1986, Animal Behav., 34: 773–784, reported on vocalizations. See brief account by Tyler and Knight, 2009, Field Guide Frogs Aust.: 148–149. See brief account, as Uperoleia capitulata, by Tyler and Knight, 2009, Field Guide Frogs Aust.: 148–149.

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.