Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nyctimystes cheesmani Tyler, 1964

Class: Amphibia > Order: Anura > Family: Pelodryadidae > Genus: Nyctimystes > Species: Nyctimystes cheesmani

Nyctimystes montana Parker, 1936, Ann. Mag. Nat. Hist., Ser. 10, 17: 80. Holotype: BMNH 1947.2.24.31 (formerly 1935.3.9.176) according to Condit, 1964, J. Ohio Herpetol. Soc., 4: 96. Type locality: "Mondo, Papua (5000 ft.)", Central Province, Papua New Guinea. Secondary homonym of Hyla montana Peters and Doria, 1878.

Nyctimystes cheesmani Tyler, 1964, Zool. Abh. Staatl. Mus. Tierkd. Dresden, 27: 268. Replacement name for Nyctimystes montana Parker, 1936.

Nyctimystes cheesmanae — Menzies, 1976, Handb. Common New Guinea Frogs: 45. Unjustified emendation according to Dubois, 2007, Zootaxa, 1550: 67, and Opinion 2447 of the ICZN (Anonymous, 2020, Bull. Zool. Nomencl., 77: 48–49). See comment.

Litoria cheesmanae — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 362. See comment.

Nyctimystes cheesmani — Kraus, 2013, Mem. Queensland Mus., 56 : 581–587, by implication; Donnellan, Mahony, Esquerré, Brennan, Price, Lemmon, Lemmon, Günther, Monis, Bertozzi, Keogh, Shea, and Richards, 2025, Zool. J. Linn. Soc., 204(2: zlaf015): 57.

Common Names

Cheesman's Big-eyed Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 60).

Distribution

Montane Central Province, Papua New Guinea.

Geographic Occurrence

Natural Resident: Papua New Guinea

Endemic: Papua New Guinea

Comment

For discussion see Zweifel, 1958, Am. Mus. Novit., 1896: 21 (as Nyctimystes montana)Menzies, 2006, Frogs New Guinea & Solomon Is.: 155–156, provided a brief account (as Nyctimystes cheesmanae) and suggested that more than one species may exist under this name. Brandon-Jones, Duckworth, Jenkins, Rylands, and Sarmiento, 2007, Zootaxa, 1541: 41–48, and Dubois, 2007, Zootaxa, 1550: 49–68, discussed the nomenclatural principles that prevent the name cheesmani from being emendable to cheesmanae. Kraus, 2012, Zootaxa, 3493: 1–26, redelimited the species and provided an account. Opinion 2447 of the ICZN (Anonymous, 2020, Bull. Zool. Nomencl., 77: 48–49: 47–48) noted that the original spelling of the species name is to be maintained. Ferreira, Oliver, Kraus, Günther, Richards, Tjaturadi, Arida, Hamidy, Riyanto, Trilaksono, Thébaud, and Fouquet, 2025, Frontiers Biogeograph., 18(e137988): 1–15, suppl. files, suggested that the nominal species is a complex on the basis of molecular and acoustic data.

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