Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Spicicalyx papua (Van Kampen, 1909)

Class: Amphibia > Order: Anura > Family: Pelodryadidae > Genus: Spicicalyx > Species: Spicicalyx papua

Hyla papua Van Kampen, 1909, Nova Guinea, 9: 33. Holotype: ZMA 5679, according to Van Tuijl, 1995, Bull. Zool. Mus. Univ. Amsterdam, 14: 127. Type locality: "Noord-Fluss, bei Bivak-Insel", Lorentz River, Southern Lowlands, New Guinea (Indonesia). A secondary homonym of Nyctimantis papua Boulenger, 1897.

Litoria papua — Richards, Hoskin, Cunningham, McDonald, and Donnellan, 2010, Zootaxa, 2391: 42. Disclaimed resurrection of name, although noted distinct species status.

Spicacalyx papua — Richards, Tjaturadi, Mumpuni, and Puradyatmika, 2015, Field Guide Mimika Region: 22. Omitted in error by Donnellan, Mahony, Esquerré, Brennan, Price, Lemmon, Lemmon, Günther, Monis, Bertozzi, Keogh, Shea, and Richards, 2025, Zool. J. Linn. Soc., 204(2: zlaf015): 1–80 (S. Richards, personal commun., June 2025).

Common Names

None noted.

Distribution

Known only from the Mimika area of the southern lowlands of Papua, New Guinea, Indonesia.

Geographic Occurrence

Natural Resident: Indonesia, Indonesia - Papua Region

Comment

Noted as being a distinct species from Litoria genimaculata by Richards, Hoskin, Cunningham, McDonald, and Donnellan, 2010, Zootaxa, 2391: 33-46, in whose synonymy it had been placed by Tyler, 1964, Zool. Abh. Staatl. Mus. Tierkd. Dresden, 27: 266. The resurrection of the species name was disclaimed pending either coining of a replacement name or, presumably, the partition of nominal Litoria. Placed in Spicicalyx after the inadvertent exclusion from the major revision of pelodryadid frogs by Donnellan, Mahony, Esquerré, Brennan, Price, Lemmon, Lemmon, Günther, Monis, Bertozzi, Keogh, Shea, and Richards, 2025, Zool. J. Linn. Soc., 204(2: zlaf015): 1–80.

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