Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Amietia fuscigula (Duméril and Bibron, 1841)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Amietia > Species: Amietia fuscigula

Rana fuscigula Duméril and Bibron, 1841, Erp. Gen., 8: 386. Holotype: MNHNP 4471, according to Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 36. See comments by Blackburn and Scali, 2014, Herpetol. Monogr., 28: 40. Type locality: "environs du cap de Bonne-Espérance", Rep. South Africa.

Rana fuscigula fuscigula — Hewitt, 1927, Rec. Albany Mus., 3: 408; Parker, 1936, Novit. Zool., 40: 141.

Rana (Rana) fuscigula — Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat.: 36; Mertens, 1971, Abh. Senckenb. Naturforsch. Ges., 529: 7; Dubois, 1987 "1986", Alytes, 5: 42.

Rana (Afrana) fuscigula — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 334.

Afrana fuscigula — Visser and Channing, 1997, J. Afr. Zool., 111: 192.

Amietia fuscigula — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 366.

English Names

Dark-throated River Frog (Hewitt, 1937, Guide Vert. Fauna E. Cape Province, Rept. Amph. Fishes: 95; Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 132).

Dusky-throated River Frog (Rose, 1950, Rep. Amph. S. Afr.: 46; Cochran, 1961, Living Amph. World: 162; Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 132).

Dark-throated Frog (Van Dijk, 1978, J. Herpetol. Assoc. Afr., 17: 15).

Cape River Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 132; Channing, 2001, Amph. Cent. S. Afr.: 259; Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 273; Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 400; Channing, Dehling, Lötters, and Ernst, 2016, Zootaxa, 4155: 27).

Cape Rana (Wager, 1965, Frogs S. Afr.: 141; Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 132).

Brown-throated Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 107).

Cape River Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 354).

Distribution

Southern parts of Rep. South Africa, from Rhodes in the east to the vicinity of Cape Town on the west.

Geographic Occurrence

Natural Resident: South Africa

Endemic: South Africa

Comment

Prior to the revision of Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297, placed in the Section Strongylopus, subgenus Afrana of Rana of Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 334. Reviewed by Poynton, 1964, Ann. Natal Mus., 17: 103–109, who clarified the Rana angolensis–Rana fuscigula problem. Further diagnosis in Channing, 1978, Ann. Natal Mus., 23: 361–365. See brief account by Lambiris, 1988, Lammergeyer, 39: 93–94, and longer ones by Channing, 2001, Amph. Cent. S. Afr.: 259–260, Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 273–274, Pickersgill, 2007, Frog Search: 96–97, and Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 400–401. The name Rana hymenopus, although properly a synonym of Rana fuscigula has generally been applied to the species currently referred to as Amietia umbraculata. Mercurio, 2011, Amph. Malawi: 283, noted an unnamed and related species in Malawi. See comment under Strongylopus hymenopus. Channing and Baptista, 2013, Zootaxa, 3640: 501–520, demonstrated broad confusion in the application of this name outside of the Cape Region of southern South Africa. Channing, Rödel, and Channing, 2012, Tadpoles of Africa: 319–320, provided information on comparative larval morphology. See account and revision by Channing, Dehling, Lötters, and Ernst, 2016, Zootaxa, 4155: 28–31. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 354–355, provided a brief account, photograph, and range map.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.