Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Natalobatrachus Hewitt and Methuen, 1912

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Natalobatrachus

1 species

Natalobatrachus Hewitt and Methuen, 1912, Trans. R. Soc. S. Afr., 3: 107. Type species: Natalobatrachus bonebergi Hewitt and Methuen, 1912, by monotypy.

Natalobatrachus — Laurent, 1941, Rev. Zool. Bot. Afr., 34: 194. Consideration as a subgenus of Phrynobatrachus.

English Names

Kloof Frogs (Passmore and Carruthers, 1979, S. Afr. Frogs: 172; Ananjeva, Borkin, Darevsky, and Orlov, 1988, Dict. Amph. Rept. Five Languages: 116).

Natal Diving Frogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 102).

Distribution

As for the single species.

Comment

Noble, 1924, Bull. Am. Mus. Nat. Hist., 49: 335; Laurent, 1941, Rev. Zool. Bot. Afr., 34: 194; Laurent, 1957, Rev. Zool. Bot. Afr., 56: 274; and Loveridge, 1957, Bull. Mus. Comp. Zool., 117: 346, included Natalobatrachus in Phrynobatrachus. See Poynton, 1964, Ann. Natal Mus., 17: 143-144 (who resurrected Natalobatrachus), and rejected the inclusion of other species of Phrynobatrachus, Poynton, 1976, Rev. Zool. Bot. Afr., 90: 218-219, and Lambiris, 1988, Lammergeyer, 39: 108-109, for accounts and discussion. Scott, 2005, Cladistics, 21: 531, suggested that Natalobatrachus is in Phrynobatrachidae rather than in Cacosterninae as hypothesized by Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 136. Wiens, Sukumaran, Pyron, and Brown, 2009, Evolution, 63: 1217-1231, provided corroboration for this taxon to rest within Cacosterninae and suggested it to be the sister taxon of Arthroleptella, a result confirmed by Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543-583, in their study of Genbank sequences.

Contained taxa (1 sp.):

Natalobatrachus bonebergi Hewitt and Methuen, 1912

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