Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tomopterna marmorata (Peters, 1854)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Tomopterna > Species: Tomopterna marmorata

Pyxicephalus marmoratus Peters, 1854, Ber. Bekannt. Verhandl. K. Preuss. Akad. Wiss. Berlin, 1854: 627. Holotype: ZMB 4808 according to Bauer, Günther, and Klipfel, 1995, in Bauer et al. (eds.), Herpetol. Contr. W.C.H. Peters: 49. Type locality: "Boror", Mozambique.

Tomopterna marmorata — Günther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 7; Vences, Glaw, Kosuch, Das, and Veith, 2000, Lourenço and Goodman (eds.), Diversité et Endéémisme à Madagascar: 232.

Pyxicephalus obtusus FitzSimons, 1930, Ann. Transvaal Mus., 14: 40. Types: TMP 12901, by original designation. Type locality: "the farm 'Zekoegat' on the Olifants River, Eastern Transvaal", Rep. South Africa. Synonymy by Pienaar, 1963, Koedoe, 6: 80; Poynton, 1964, Ann. Natal Mus., 17: 99.

Arthroleptis rosei Hoffman, 1944, Soöl. Navors. Nas. Mus. Bloemfontein, 1: 174. Holotype: NMP 2614, by original designation. Type locality: "Chitiala [= Chitala River], Nyasaland", Malawi. Synonymy with Rana delalandii by Loveridge, 1953, Bull. Mus. Comp. Zool., 110: 376, and with Tomopterna marmorata by Poynton, 1964, Ann. Natal Mus., 17: 99.

Pyxicephalus marmoratus — Poynton, 1964, Ann. Natal Mus., 17: 99. Removal from synonymy of Tomopterna delalandii.

Rana (Tomopterna) marmorata — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Tomopterna (Tomopterna) marmorata — Dubois, 1987 "1986", Alytes, 5: 57.

Common Names

Striped Pyxie (Wager, 1965, Frogs S. Afr.: 134).

Blunt-nosed Pyxie (Pienaar, 1963, Koedoe, 6: 80).

Blunt-nosed Burrowing Frog (Pienaar, 1963, Koedoe, 6: 80).

Mozambique Pyxie (Broadley, 1971, Puku, 6: 112).

Mozambique Dwarf Bullfrog (Broadley, 1974, J. Herpetol. Assoc. Afr., 12: 31).

Ruddy Burrowing Frog (Broadley, 1978, J. Herpetol. Assoc. Afr., 19: 33).

Russet-backed Sand Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 124; Channing, 2004, in Minter et al. (eds.), Atlas Frogs S. Afr. Lesotho and Swaziland: 325; Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 442).

Marmorate Pyxie (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5; Passmore and Carruthers, 1979, S. Afr. Frogs: 124; Lambiris, 1990 "1989", Monogr. Mus. Reg. Sci. Nat. Torino, 10: 87).

Mozambique Burrowing Frog (Broadley, 1973, J. Herpetol. Assoc. Afr., 10: 23; Passmore and Carruthers, 1979, S. Afr. Frogs: 124).

Moçambique Burrowing Frog (Passmore and Carruthers, 1978, J. Herpetol. Assoc. Afr., 19: 5).

Marbled Bullfrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Marbled Sand Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 380).

Distribution

Northeastern Rep. South Africa, and Mozambique through Malawi to eastern Botswana and southern Zambia; Kenya highlands, possibly to be found in intervening Tanzania and possibly into Eswatini.

Geographic Occurrence

Natural Resident: Botswana, Kenya, Malawi, Mozambique, South Africa, Zambia, Zimbabwe

Likely/Controversially Present: Eswatini, Tanzania

Comment

Possibly has a wider distribution northward than indicated but obscured by confusion with other species, particularly Tomopterna cryptotis, in the literature according to J.C. Poynton In Inger, 1985, in Frost (ed.), Amph. Species World: 524. Auerbach, 1987, Amph. Rept. Botswana: 46, provided a summary account and grid map for Botswana. See accounts in Poynton and Broadley, 1985, Ann. Natal Mus., 27: 129–130, Lambiris, 1988, Lammergeyer, 39: 78–79, Pickersgill, 2007, Frog Search: 122–123, and Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 442–443. Mercurio, 2011, Amph. Malawi: 294–295, provided an account for Malawi. Du Preez and Carruthers, 2017, Frogs S. Afr., Compl. Guide: 470–471, provided an account, including a polygon range map, photograph, identification features, adult morphology, habitat, and call. See Bittencourt-Silva, 2019, Amph. Rept. Conserv., 13 (2: e181): 12–13, for comment on population in western Zambia. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 380–381, provided a brief account, photograph, and range map.

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist