Tomopterna milletihorsini (Angel, 1922)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Tomopterna > Species: Tomopterna milletihorsini

Arthroleptis milleti-horsini Angel, 1922, Bull. Mus. Natl. Hist. Nat. Paris, 28: 41. Holotype: MNHNP 1921.635 by original designation; not mentioned by Guibé, 1950 "1948", Cat. Types Amph. Mus. Natl. Hist. Nat., and regarded as lost by Ohler and Frétey, 2008, Alytes, 25: 173–175. Type locality: "la région caillouteuse de Bélédougou, à Kati (12 kilomètres au nord de Bamakko)", Mali.

Arthroleptis agadezi Angel, 1936, Bull. Soc. Zool. France, 61: 275. Syntypes: MNHNP 36.124–125 (2 specimens), according to the original publication. Type locality: "Agadez (Aïr)", Niger. Considered a junior synonym of Pyxicephalus delalandii (Tschudi) (Somali specimens of this nominal species) by Angel and Guibé, 1948, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 20: 62, before the firm distinction of Tomopterna cryptotis and Tomopterna delalandii. Treated by implication as a synonym of Tomopterna milletihorsini by Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 397.

Schoutedenella milletihorsiniLaurent, 1954, Ann. Mus. R. Congo Belge, Tervuren, Ser. Quarto, Sci. Zool., 1: 40; Laurent, 1985, in Frost (ed.), Amph. Species World: 19.

Arthroleptis milletihorsiniLaurent, 1957, Rev. Zool. Bot. Afr., 56: 274–275, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 336.

Tomopterna milletihorsiniOhler and Frétey, 2008, Alytes, 25: 174.

English Names

Mali Screeching Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 37).

Horsin's Sand Frog (Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 382).


North and East subsaharan Africa from southern Algeria, Mauritania, and Guinea east through northern Nigeria, northern Cameroon, Burkina Faso, northern Central African Rep., southern Sudan and South Sudan to the Ethiopian border, thence south through Kenya and northern Tanzania. 

Geographic Occurrence

Natural Resident: Algeria, Burkina Faso, Cameroon, Central African Republic, Egypt, Guinea, Kenya, Mali, Mauritania, Niger, Nigeria, Senegal, South Sudan, Sudan, Tanzania


See comments under Tomopterna cryptotis for possibly relevant literature. Schiøtz, 1963, Vidensk. Medd. Dansk Naturhist. Foren., 125: 30, provided records for Nigeria (as Tomopterna delalandii). See Salvador, 1996, Smithson. Herpetol. Inform. Serv., 109,: 36– for account (as Tompterna cryptotis) for Mauritania and Niger. Rödel, 2000, Herpetofauna W. Afr., 1: 76–78, provided an account (as Tomopterna cryptotis). Padial, 2003, Herpetol. Rev., 34: 260, reported the species (as Tomopterna cryptotis) for Mauritania. Padial and De la Riva, 2004, Rev. Esp. Herpetol., 18: 95, provided records from Mauritania (as Tomopterna cryptotis). Ohler and Frétey, 2008, Alytes, 25: 173–175, discussed the identity of the type and the type locality. Zimkus and Larson, 2011, Zootaxa, 2933: 27–45, provided the Nara, Mali record. Padial, Crochet, Géniez, and Brito, 2013, Basic & Appl. Herpetol., 27: 11–22, discussed systematics (as Tomopterna cryptotis), distribution, and conservation status in Mauritania.  Ohler and Frétey, 2014, J. E. Afr. Nat. Hist., 103: 91, provided a brief discussion of a collection from northern Mozambique. Sow, Gonçalves, Sousa, Martínez-Freiría, Santarém, Velo-Antón, Dieng, Campos, Diagne, Boratyński, and Brito, 2017, Basic & Appl. Herpetol., 31: 101–116, reported the species from extreme southeastern Mauritania. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 382–383, provided a brief account, photograph, and range map, as Tomopterna cf. milletihorsini. See account for Mauritania by Escoriza and Ben Hassine, 2019, Amph. N. Afr.: 281–284. Spawls, Wasonga, and Drewes, 2019, Amph. Kenya: 35, provided a range map for Kenya, photograph, and brief characterization, as Tomopterna cryptotis. Ayoro, Segniagbeto, Hema, Penner, Oueda, Dubois, Rödel, Kabré, and Ohler, 2020, Zoosystema, 42: 547–582, discussed records, identification, and habitat in Burkina Faso. Sampaio, Velo-Antón, Martínez-Freiría, Sánchez-Vialas, Pleguezuelos, Géniez, Crochet, and Brito, 2021, Conserv. Genetics, 22: 233–248, discussed landscape genetics and range in Mauritania. Pagano, Daf, Ballouche, Taïbi, and Sinsch, 2022, Salamandra, 58: 317–322, reported on the advertisement call from  Diawling National Park, southwestern Mauritania. 

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