Tomopterna tuberculosa (Boulenger, 1882)

Class: Amphibia > Order: Anura > Family: Pyxicephalidae > Subfamily: Cacosterninae > Genus: Tomopterna > Species: Tomopterna tuberculosa

Pyxicephalus rugosus Günther, 1865 "1864", Proc. Zool. Soc. London, 1864: 479. Syntypes: BMNH (2 specimens), according to XXX. Type locality: "Pungo Andongo", Angola, and "W. Africa"; specimens noted from "W. Africa" by Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 30, were not considered types by him. Preoccupied by Rana rugosa Temminck and Schlegel, 1838.

Rana tuberculosa Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 30. Replacement name for Pyxicephalus rugosus Günther, 1864.

Rana pulchra Boulenger, 1896, Ann. Mag. Nat. Hist., Ser. 6, 18: 468. Holotype: BMNH, by original designation. Type locality: "Lake Tanganyika". Synonymy by Loveridge, 1957, Bull. Mus. Comp. Zool., 117: 345 (with Rana tuberculosa); Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 36.

Rana (Tomopterna) signata Ahl, 1927 "1925", Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1925: 43. Syntypes: ZMB (2 specimens), unnumbered in original publication. Type locality: "Huilla", Angola. Secondary homonym of Polypedates signata Günther, 1872 (= Rana signata). Synonymy by Laurent, 1954, Publ. Cult. Companhia Diamantes Angola, 23: 72.

Rana (Tomopterna) cacondana Ahl, 1927 "1925", Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1925: 44. Holotype: ZMB 15477, by museum records; unnumbered in original publication. Type locality: "Caconda", Angola. Synonymy by Laurent, 1954, Publ. Cult. Companhia Diamantes Angola, 23: 72.

Rana (Tomopterna) pulchraDe Witte, 1930, Rev. Zool. Bot. Afr., 19: 263.

Pyxicephalus tuberculosusParker, 1936, Novit. Zool., 40: 142. Poynton, 1964, Ann. Natal Mus., 17: 100.

Rana (Tomopterna) tuberculosaMonard, 1937 "1936", Bull. Soc. Neuchatel. Sci. Nat., 62: 44.

Tomopterna cacondaDeckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 142.

Tomopterna rugosaLaurent, 1954, Publ. Cult. Companhia Diamantes Angola, 23: 72.

Tomopterna tuberculosaLaurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 132. Poynton and Broadley, 1985, Ann. Natal Mus., 27: 130-131; Vences, Glaw, Kosuch, Das, and Veith, 2000, Lourenço and Goodman (eds.), Diversité et Endéémisme à Madagascar: 232.

Rana (Tomopterna) tuberculosaDubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 233, by implication.

Tomopterna (Tomopterna) tuberculosaDubois, 1987 "1986", Alytes, 5: 57.

English Names

Warty Frog (Cochran, 1961, Living Amph. World: 158).

Beaded Pyxie (Broadley, 1971, Puku, 6: 113; Lambiris, 1990 "1989", Monogr. Mus. Reg. Sci. Nat. Torino, 10: 89).

Beaded Burrowing Frog (Broadley, 1973, J. Herpetol. Assoc. Afr., 10: 23).

Beaded Dwarf Bullfrog (Broadley, 1974, J. Herpetol. Assoc. Afr., 12: 31).

Angola Bullfrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Rough Sand Frog (Channing and Howell, 2006, Amph. E. Afr.: 333; Du Preez and Carruthers, 2017, Frogs S. Afr., Compl. Guide: 476).

Tuberculate Sand Frog (Channing and Howell, 2006, Amph. E. Afr.: 333 [alternative name]).

Bearded Pyxie (Channing and Howell, 2006, Amph. E. Afr.: 333 [alternative name]).

Bearded Sand Frog (Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 448).

Distribution

Angola and northwestern Namibia to Tanzania and Zimbabwe.

Comment

See Laurent, 1964, Publ. Cult. Companhia Diamantes Angola, 67: 133, for synonymy. See accounts by Poynton and Broadley, 1985, Ann. Natal Mus., 27: 130-131, Channing and Howell, 2006, Amph. E. Afr.: 333-334, Pickersgill, 2007, Frog Search: 124-125, and Du Preez and Carruthers, 2009, Compl. Guide Frogs S. Afr.: 448-449. See also (as Rana tuberculosa) Schmidt and Inger, 1959, Explor. Parc Natl. Upemba, Miss. G.F. de Witte, 56: 36-38. Dawood, Channing, and Bogart, 2002, Mol. Phylogenet. Evol., 22: 407-413, reported two unnamed species masquerading under this name from Beira, Mozambique, and Shankara, Namibia. The Shankara specimen was subsequently confirmed as Tomopterna cryptotis by Channing and Du Preez, 2020, Alytes, 37: 1–21. Mercurio, 2011, Amph. Malawi: 296-297, provided an account for Malawi. Marques, Ceríaco, Blackburn, and Bauer, 2018, Proc. California Acad. Sci., Ser. 4, 65 (Suppl. II): 155–156, provided a map for Angola and brief discussion of the literature. See Wilson and Channing, 2019, Zootaxa, 4609: 225–246, who provided a range map. Butler, Ceríaco, Marques, Bandeira, Júlio, Heinicke, and Bauer, 2019, Herpetol. Rev., 50: 225–240, reported on a collection from Huila Province, southwestern Angola. Channing and Rödel, 2019, Field Guide Frogs & Other Amph. Afr.: 382–383, provided a brief account, photograph, and range map. 

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