Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylarana megalonesa (Inger, Stuart, and Iskandar, 2009)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Hylarana > Species: Hylarana megalonesa

Rana megalonesa Inger, Stuart, and Iskandar, 2009, Zool. J. Linn. Soc., 155: 136. Holotype: FMNH 267821, by original designation. Type locality: "Bukit Sarang (2°39′N/113°03′E), Bintulu Division, Sarawak (Borneo), Malaysia".

Chalcorana megalonesa — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 33. See comment under Ranidae record.

Chalcorana megalonesa — Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 187.

Hylarana megalonesa — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 423.

Hylarana (Chalcorana) megalonesa — Wu, Xu, Suwannapoom, Nguyen, Murphy, Papenfuss, Lathrop, Kilunda, Gao, Yuan, Chen, Zhang, Zhao, Wang, Rahman, Nneji, Zhao, Wang, Jin, Zhang, and Che, 2024 "2025", Mol. Phylogenet. Evol., 108223: 1–11, by implication.

Common Names

Large White-lipped Frog (Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 327).

Distribution

Western and northern Borneo (Sarawak and Sabah, Malaysia, likely into adjacent Kalimantan, Indonesia); Java, Indonesia.

Geographic Occurrence

Natural Resident: Indonesia, Malaysia, Malaysia, East (Sarawak and/or Sabah)

Comment

Confused with Hylarana chalconota prior to its naming according to the original publication. Malkmus, Manthey, Vogel, Hoffmann, and Kosuch, 2002, Amph. Rept. Mount Kinabalu: 159-160, provided an account for Rana chalconota that presumably applies to this species. Inger and Stuebing, 2009, Raffles Bull. Zool., 57: 534, commented on the range. Haas, Kueh, Joseph, bin Asri, Das, Hagmann, Schwander, and Hertwig, 2018, Evol. Syst., 2: 89–114, provided a brief account of morphology and natural history for the Sabah population. Gillespie, Ahmad, and Shia, 2021, Field Guide Frog Lower Kinabatangan Region Sabah: 31, provided a brief account, summarizing identification and life history in the Lower Kinabatangan Region, Sabah, Malaysia. Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 327–329, summarized the knowledge of habitat, reproduction, larval morphology and coloration. Sabinhaliduna, Arisuryanti, and Hamidy, 2022, Treubia, 49: 97–114, provided a genetically confirmed record from Pasir Angin, Bogor (Java), Indonesia, where it occurs in sympatric with Chalcorana chalconota. Al Gifari, Hasyalillah, Albiansyah, Mawla, Bainahum, Rozaq, Al Ghiyasi, Murfi, Fitriana, and Haribowo, 2026, J. Biol. Tropis, 26: 607–616, reported this species at Tapos Resort, Gunung Pangrango National Park, western Java, Indonesia.

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For additional information see AmphibiaWeb report
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For related information on conservation and images as well as observations see iNaturalist