Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Clinotarsus penelope Grosjean, Bordoloi, Chuaynkern, Chakravarty, and Ohler, 2015

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Clinotarsus > Species: Clinotarsus penelope

Clinotarsus penelope Grosjean, Bordoloi, Chuaynkern, Chakravarty, and Ohler, 2015, Zootaxa, 4058: 484. Holotype: Tadpole MNHNP 2000.4630 in stage 36, by original designation. Type locality: "Namtok Raman Forest Park (8°27’ N, 98°32’ E, WGS 84, about 30 m asl), Phang Nga Province, Thailand".

Common Names

Hill Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 140).

Palebrown Stream Frog (Nutphund, 2001, Amph. Thailand: 121).

Penelope's Tadpole Frog (Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 54.

Phang Nga Hill Frog (Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 46).

Malay Pointed-snout Frog (Zug, 2022, Smithson. Contrib. Zool., 653: 36).

Distribution

Peninsular Thailand from central Kachana Buri province south to Trang Province, and adjacent peninsular Myanmar (Mon and Tanintharyi).

Geographic Occurrence

Natural Resident: Myanmar, Thailand

Comment

With the naming of Clinotarsus penelope, based on larval characters, the literature of Clinotarsus alticola from Thailand must be presumed to apply to the newer species. See account by Taylor, 1962, Univ. Kansas Sci. Bull., 43: 453-456 (as Rana alticola). Chan-ard, 2003, Photograph. Guide Amph. Thailand: 140-141, provided a very brief account, map for Thailand, and photograph, as Rana cf. alticola. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 84-86, to compare this species to other ranids of the Sunda Shelf region, although this species was conflated with Clinotarsus alticola. In the original publication, the authors provided molecular evidence that possibly one more species of Clinotarsus will be named from the Tak-Kanchanaburi Province boundary area of Thailand. Chan-ard, Cota, and Makchai, 2011, Amph. E. Region Thailand: 142, detailed the range (as Clinotarsus alticola) in Thailand. Mulcahy, Lee, Miller, Chand, Thura, and Zug, 2018, ZooKeys, 757: 95, provided a record (as Clinotarsus alticola) from Taninthary Division, southern Myanmar. Zug, Mulcahy, Vindum, and Wilkinson, 2018, Alytes, 36: 270, provided a record for South Tanintharyi, peninsular Myanmar. Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 54, provided a brief account for South Tanintharyi, peninsular Myanmar. Zug, 2022, Smithson. Contrib. Zool., 653: 36, briefly discussed identification, habitat, and range in Myanmar.

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For information on distribution, habitat, and conservation see the Map of Life
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