Humerana humeralis (Boulenger, 1887)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Humerana > Species: Humerana humeralis

Rana humeralis Boulenger, 1887, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 5: 420. Syntypes: MSNG (several specimens) and BMNH 1947.2.3.33–45 (originally numbered 1889.3.25.45–46), presumably on exchange from MSNG, according to museum records; MSNG 29299 designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 215. Type locality: "Bhamò, [and] Teinzò", Myanmar; restricted to Bhamò, Myanmar, by lectotype designation. Biju, Garg, Gokulakrishnan, Sivaperuman, Thammachoti, Ren, Gopika, Bisht, Hamidy, and Shouche, 2020, Zootaxa, 4878: 31, discussed the location of Bhamò. 

Rana (Hylorana) humeralisBoulenger, 1920, Rec. Indian Mus., 20: 124.

Hylorana humeralisDeckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 144.

Hylarana humeralis —  Bourret, 1942, Batr. Indochine: 98; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 496, by implication. 

Rana (Hylarana) humeralisBourret, 1942, Batr. Indochine: 324. Dubois, 1987 "1986", Alytes, 5: 42; by implication.

Rana (Humerana) humeralisDubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 324-325.

Humerana humeralisFrost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 35. See comment under Ranidae record.

English Names

Bhamo Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 107; Ahmed, Das, and Dutta, 2009, Amph. Rept. NE India: 42; Dinesh, Radhakrishnan, Gururaja, and Bhatta, 2009, Rec. Zool. Surv. India, Occas. Pap., 302: 73).

Bhamò Frog (Schleich, Anders, and Kästle, 2002, in Schleich and Kästle (eds.), Amph. Rept. Nepal: 80).

Boulenger's Green Frog (Mathew and Sen, 2010, Pict. Guide Amph. NE India: 77).

Northern Caped Frog (Zug, 2022, Smithson. Contrib. Zool., 653: 36).


Northern Myanmar (Kachin and Sagaing) through northeastern India (Arunachal Pradesh, Assam, and northernmost West Bengal) to southeastern Nepal and eastern Bangladesh at low elevations; presumably in intermediate regions of Bhutan.


See accounts by Boulenger, 1920, Rec. Indian Mus., 20: 148; Bourret, 1942, Batr. Indochine: 324-325. Anders, 2002, in Schleich and Kästle (eds.), Amph. Rept. Nepal: 293-295, provided an account for the Nepal population. Ao, Bordoloi, and Ohler, 2003, Zoos' Print J., 18: 1117-1125, presented a specific locality for Nagaland, northeastern India. Ningombam and Bordoloi, 2007, Zoos' Print J., 22: 2688-2690, provided a record for Manipur, northeastern India. Hussain, Gogoi, Choudhury, Baruah, Sharma, Kalita, and Sengupta, 2007, Russ. J. Herpetol., 14: 167-171, provided records (as Rana humeralis) for Assam and Arunachal Pradesh, northeastern India, and provided morphometric and natural history information. Sen and Mathew, 2009, Rec. Zool. Surv. India, Occas. Pap., 285: 123, provided a specific record for Nagaland, northeastern India. Das and Dutta, 2007, Hamadryad, 31: 154–181, noted no larval descriptions in the literature. Ahmed, Das, and Dutta, 2009, Amph. Rept. NE India: 42, provided a brief account for northeastern India. Mathew and Sen, 2010, Pict. Guide Amph. NE India: 77, provided a brief characterization and photograph. See Shah and Tiwari, 2004, Herpetofauna Nepal: 70, for brief account (as Rana humeralis). Bortamuli, Bordoloi, Ohler, and Grosjean, 2010, J. Nat. Hist., London, 44: 421–445, described external and buccopharyngeal anatomy. Deuti, Ray, Bag, and Dey, 2017, Rec. Zool. Surv. India, 117: 221–241, provided records from the "duars region", West Bengal adjacent to Bhutan. Khatiwada, Wang, Zhao, Xie, and Jiang, 2021, Asian Herpetol. Res., 12: 1–35, did not mention this species as part of the Nepal fauna, a curious lapse. Zug, 2022, Smithson. Contrib. Zool., 653: 36, briefly discussed identification, habitat, and range in Myanmar. 

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