Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Humerana humeralis (Boulenger, 1887)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Humerana > Species: Humerana humeralis

Rana humeralis Boulenger, 1887, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 5: 420. Syntypes: MSNG (several specimens); MSNG 29299 designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 215. Type locality: "Bhamò, [and] Teinzò", Myanmar; restricted to Bhamò, Myanmar, by lectotype designation. Biju, Garg, Gokulakrishnan, Chandrakasan, Thammachoti, Ren, Gopika, Bisht, Hamidy, and Shouche, 2020, Zootaxa, 4878: 31, discussed the location of Bhamò.

Rana (Hylorana) humeralis — Boulenger, 1920, Rec. Indian Mus., 20: 124.

Hylorana humeralis — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 144.

Rana (Hylarana) humeralis — Bourret, 1942, Batr. Indochine: 324. Dubois, 1987 "1986", Alytes, 5: 42; by implication.

Rana (Humerana) humeralis — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 324-325.

Humerana humeralis — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 368. Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 35. See comment under Ranidae record.

Hylarana humeralis — Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 496, by implication.

English Names

Bhamo Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 107; Ahmed, Das, and Dutta, 2009, Amph. Rept. NE India: 42; Dinesh, Radhakrishnan, Gururaja, and Bhatta, 2009, Rec. Zool. Surv. India, Occas. Pap., 302: 73).

Bhamò Frog (Schleich, Anders, and Kästle, 2002, in Schleich and Kästle (eds.), Amph. Rept. Nepal: 80).

Boulenger's Green Frog (Mathew and Sen, 2010, Pict. Guide Amph. NE India: 77).

Distribution

Northern Myanmar through northeastern India (Arunachal Pradesh, Assam, and northernmost West Bengal) to southeastern Nepal and eastern Bangladesh at low elevations; presumably in intermediate regions of Bhutan.

Comment

See accounts by Boulenger, 1920, Rec. Indian Mus., 20: 148; Bourret, 1942, Batr. Indochine: 324-325. Anders, 2002, in Schleich and Kästle (eds.), Amph. Rept. Nepal: 293-295, provided an account for the Nepal population. Ao, Bordoloi, and Ohler, 2003, Zoos' Print J., 18: 1117-1125, presented a specific locality for Nagaland, northeastern India. Ningombam and Bordoloi, 2007, Zoos' Print J., 22: 2688-2690, provided a record for Manipur, northeastern India. Hussain, Gogoi, Choudhury, Baruah, Sharma, Kalita, and Sengupta, 2007, Russ. J. Herpetol., 14: 167-171, provided records (as Rana humeralis) for Assam and Arunachal Pradesh, northeastern India, and provided morphometric and natural history information. Sen and Mathew, 2009, Rec. Zool. Surv. India, Occas. Pap., 285: 123, provided a specific record for Nagaland, northeastern India. Das and Dutta, 2007, Hamadryad, 31: 154–181, noted no larval descriptions in the literature. Ahmed, Das, and Dutta, 2009, Amph. Rept. NE India: 42, provided a brief account for northeastern India. Mathew and Sen, 2010, Pict. Guide Amph. NE India: 77, provided a brief characterization and photograph. See Shah and Tiwari, 2004, Herpetofauna Nepal: 70, for brief account (as Rana humeralis). Bortamuli, Bordoloi, Ohler, and Grosjean, 2010, J. Nat. Hist., London, 44: 421–445, described external and buccopharyngeal anatomy. Deuti, Ray, Bag, and Dey, 2017, Rec. Zool. Surv. India, 117: 221–241, provided records from the "duars region", West Bengal adjacent to Bhutan. Khatiwada, Wang, Zhao, Xie, and Jiang, 2021, Asian Herpetol. Res., 12: 1–35, did not mention this species as part of the Nepal fauna, a curious lapse.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.