Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rana ornativentris Werner, 1903

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Rana > Species: Rana ornativentris

Rana japonica var. ornativentris Werner, 1903, Abh. Math. Physik. Cl. Bayer. Akad. Wiss., 22: 383. Holotype: ZSM 96/0 according to Glaw and Franzen, 2006, Spixiana, München, 29: 186. Type locality: "Nikko auf Nippon"; given as Akanuma and Tadenuma, Nikko [Honshu, Japan] by Stejneger, 1907, Bull. U.S. Natl. Mus., 58: 108.

Rana temporaria var. montana Okada and Kawano, 1923, Zool. Mag., Tokyo, 35: 364-366. Type(s): XXX. Type locality: XXX. Synonymy by Okada, 1928, Chosen Nat. Hist. Soc. J., 6: 30.

Rana temporaria ornativentris — Stejneger, 1924, Proc. Biol. Soc. Washington, 37: 76. Okada, 1927, Copeia, 158: 165; Okada, 1928, Chosen Nat. Hist. Soc. J., 6: 30; Sato and Takashima, 1955, Misc. Rep. Yamashina Inst. Ornithol. Zool., 6: 260-268.

Rana ornativentris — Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 191. Tomasik, 1967, Przeglad Zoologiczny, 11: 367.

Rana (Rana) temporaria ornativentris — Nakamura and Ueno, 1963, Japan. Rept. Amph. Color: 42.

Rana (Rana) ornativentris — Dubois, 1987 "1986", Alytes, 5: 41-42, by implication; Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 333; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

Rana (Laurasiarana) ornativentris — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 311, by implication; see Dubois, 2006, Mol. Phylogenet. Evol., 42: 317-330, and Hillis, 2007, Mol. Phylogenet. Evol., 42: 331-338, for discussion.

English Names

Nikko Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 109).

Montane Brown Frog (Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 79).

Distribution

Honshu, Shikoku, and Kyushu Is. as well as some outlying islands, Japan.

Geographic Occurrence

Natural Resident: Japan

Endemic: Japan

Comment

See Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 181-193, and Okada, 1966, Fauna Japon., Anura: 81-90, for review. In the Rana chensinensis group, subgenus Rana, Section Rana. Considered a synonym of Rana dybowskii (as Rana temporaria dybowskii) by Shannon, 1956, Herpetologica, 12: 38. Tanaka, Matsui, and Takenaka, 1994, Zool. Sci., Tokyo, 11: 753-757, discussed the relationships of this species with Rana japonica, Rana pirica, Rana tagoi, and Rana sakuraii. See Tanaka-Ueno, Matsui, Sato, Takenaka, and Takenaka, 1998, Japan. J. Herpetol., 17: 91-97, for phylogenetic relationships and discussion. See also Sumida and Nishioka, 1996, Zool. Sci., Tokyo, 13: 1-12, for geographical genetic divergence. Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 76-79, provided an account. Goris and Maeda, 2004, Guide Amph. Rept. Japan: 82-84, provided an account, map, and photograph.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.