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Rana ornativentris Werner, 1903
Rana japonica var. ornativentris Werner, 1903, Abh. Math. Physik. Cl. Bayer. Akad. Wiss., 22: 383. Holotype: ZSM 96/0 according to Glaw and Franzen, 2006, Spixiana, München, 29: 186. Type locality: "Nikko auf Nippon"; given as Akanuma and Tadenuma, Nikko [Honshu, Japan] by Stejneger, 1907, Bull. U.S. Natl. Mus., 58: 108.
Rana temporaria var. montana Okada and Kawano, 1923, Zool. Mag., Tokyo, 35: 364-366. Type(s): XXX. Type locality: XXX. Synonymy by Okada, 1928, Chosen Nat. Hist. Soc. J., 6: 30.
Rana temporaria ornativentris — Stejneger, 1924, Proc. Biol. Soc. Washington, 37: 76. Okada, 1927, Copeia, 158: 165; Okada, 1928, Chosen Nat. Hist. Soc. J., 6: 30; Sato and Takashima, 1955, Misc. Rep. Yamashina Inst. Ornithol. Zool., 6: 260-268.
Rana ornativentris — Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 191. Tomasik, 1967, Przeglad Zoologiczny, 11: 367.
Rana (Rana) temporaria ornativentris — Nakamura and Ueno, 1963, Japan. Rept. Amph. Color: 42.
Rana (Rana) ornativentris — Dubois, 1987 "1986", Alytes, 5: 41-42, by implication; Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 333; Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.
Rana (Laurasiarana) ornativentris — Hillis and Wilcox, 2005, Mol. Phylogenet. Evol., 34: 311, by implication; see Dubois, 2006, Mol. Phylogenet. Evol., 42: 317-330, and Hillis, 2007, Mol. Phylogenet. Evol., 42: 331-338, for discussion.
Common Names
Nikko Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 109).
Montane Brown Frog (Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 79).
Distribution
Honshu, Shikoku, and Kyushu Is. as well as some outlying islands, Japan.
Geographic Occurrence
Natural Resident: Japan
Endemic: Japan
Comment
See Kawamura, 1962, J. Sci. Hiroshima Univ., B—Zool., 20: 181-193, and Okada, 1966, Fauna Japon., Anura: 81-90, for review. In the Rana chensinensis group, subgenus Rana, Section Rana. Considered a synonym of Rana dybowskii (as Rana temporaria dybowskii) by Shannon, 1956, Herpetologica, 12: 38. Tanaka, Matsui, and Takenaka, 1994, Zool. Sci., Tokyo, 11: 753-757, discussed the relationships of this species with Rana japonica, Rana pirica, Rana tagoi, and Rana sakuraii. See Tanaka-Ueno, Matsui, Sato, Takenaka, and Takenaka, 1998, Japan. J. Herpetol., 17: 91-97, for phylogenetic relationships and discussion. See also Sumida and Nishioka, 1996, Zool. Sci., Tokyo, 13: 1-12, for geographical genetic divergence. Maeda and Matsui, 1990, Frogs Toads Japan, Ed. 2: 76-79, provided an account. Goris and Maeda, 2004, Guide Amph. Rept. Japan: 82-84, provided an account, map, and photograph.
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