Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Staurois natator (Günther, 1858)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Staurois > Species: Staurois natator

Ixalus natator Günther, 1858, Arch. Naturgesch., 24: 324. Syntypes: BMNH (3 specimens) according to Günther, 1859 "1858", Cat. Batr. Sal. Coll. Brit. Mus.: 75, these being BM 1933.9.10.9–11 according to museum records, these now renumbered 1947.2.2.93–94 and 1947.2.4.42. Type locality: "Philippinen"; restricted to "Mindanao" or "Leyte", Philippines, by Inger, 1954, Fieldiana, Zool., 33: 338.

Staurois natator — Cope, 1865, Nat. Hist. Rev., N.S., 5: 117.

Rana natatrix — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 71.

Ixalus granulatus Boettger, 1888, Ber. Offenbach. Ver. Naturkd., 1888: 57. Holotype: SMF 7002 according to Mertens, 1967, Senckenb. Biol., 48(A): 44. Type locality: "Siam"; corrected to Philippines by Boettger, 1892, Kat. Batr. Samml. Mus. Senckenb. Naturforsch. Ges.: 16. Synonymy by Ohler, 1996, Copeia, 1996: 1010-1012, who noted that this name is a secondary homonym of Rana granulatus (Boettger, 1883).

Rhacophorus granulosus Ahl, 1927, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1927: 38. Replacement name for Ixalus granulatus Boettger, 1888 "(Präocc. durch Rana granulatus Boettger, 1884)" in Rhacophorus.

Common Names

Mindanao Rock Frog (Diesmos, del Prado, Kim, Caguimbal, Venturina, Lorenzo, and Diesmos, 2026, ZooKeys, 1269: 309).

Mindanao Splash Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Distribution

Mindanao, Leyte, and Samar Islands Philippines (see comment).

Geographic Occurrence

Natural Resident: Philippines

Endemic: Philippines

Comment

Discussed by Inger, 1954, Fieldiana, Zool., 33: 335–344. See Ohler, 1996, Copeia, 1996: 1010–1012, for nomenclatural discussion of several of the names in synonymy. See identification table (as including Staurois nubilus) by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 84–86, to compare this species to other ranids of the Sunda Shelf region. Malkmus, Manthey, Vogel, Hoffmann, and Kosuch, 2002, Amph. Rept. Mount Kinabalu: 171–172, provided an account. Arifin, Iskandar, Bickford, Brown, Meier, and Kutty, 2011, Zootaxa, 2744: 39–52, implied that the populations of Leyte and Samar islands, when sampled genetically, may be found to be distinct species. See brief notes for Mindanao, Philippines, by Sanguila, Cobb, Siler, Diesmos, Alcala, and Brown, 2016, ZooKeys, 624: 42–44. Rosales, Yu, and Nanual, 2023, Davao Res. J., 14: 50–65, reported on distribution and habitat with special reference to the Mt. Hamiguitan Range Wildlife Sanctuary and surrounding areas of Mindanao. Flores, Mondejar, Caballero, Maceda, Olowa, and Nuñeza, 2025, Biodiversitas, 26: 3769–3780, discussed the habitat in Mount Gutom Protected Landscape on Zamboanga del Norte, Mindanao, Philippines. Diesmos, del Prado, Kim, Caguimbal, Venturina, Lorenzo, and Diesmos, 2026, ZooKeys, 1269: 303–328, discussed the range on Samar I., Philippines. Sanguila, Labaja, Bernstein, and Brown, 2026, ZooKeys, 1272: 203–251, reported the species from the Agusan Marsh Wildlife Sanctuary, eastern Mindanao I., Philippines. Baguio and Nuñeza, 2026, Internatl. J. Multidiscipl. Res., 8: 1–19, discussed habitat preference in the Mt. Sumagaya Range, Misamis Oriental, Mindanao, Philippines.

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For additional information see AmphibiaWeb report
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For related information on conservation and images as well as observations see iNaturalist