Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nyctixalus anodon (Van Kampen, 1907)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Nyctixalus > Species: Nyctixalus anodon

Rhacophorus anodon Van Kampen, 1907, Zool. Ergebn. Reis. Niederland. Ost-Indien, 4: 400. Holotype: ZMA 5707, according to Van Tuijl, 1995, Bull. Zool. Mus. Univ. Amsterdam, 14: 129. Type locality: "Schlucht bei Ajer Mantjur (Kaju tanam)" = cleft near Ajer Mantjur, Kaju tanam, Sumatra, Indonesia.

Philautus anodon — Van Kampen, 1923, Amph. Indo-Austral. Arch.: 269.

Rhacophorus (Philautus) anodon — Ahl, 1931, Das Tierreich, 55: 68.

Hazelia anodon — Liem, 1970, Fieldiana, Zool., 57: 96.

Nyctixalus anodon — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257; Delorme, Dubois, Grosjean, and Ohler, 2005, Bull. Mens. Soc. Linn. Lyon, 74: 166; Ibnudarda, Kadafi, Priambodo, Farajallah, Perwitasari-Farajallah, and Hamidy, 2025, J. Trop. Biodiversity Biotechnol., 10 (jtbb13642): 1; Dever, McGuire, Iskandar, Hamidy, and Grismer, 2026, Zootaxa, 5777: 283.

Common Names

Sumatra Indonesian Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 111).

Distribution

Sumatra and nearby islands to the west, Indonesia.

Geographic Occurrence

Natural Resident: Indonesia

Endemic: Indonesia

Comment

Synonymy (with Ixalus pictus) by Smith, 1931, Bull. Raffles Mus., 5: 19, and Inger, 1966, Fieldiana, Zool., 52: 349–350. This synonymy provisionally not accepted pending revisionary studies by Bossuyt and Dubois, 2001, Zeylanica, 6: 39. Ibnudarda, Kadafi, Priambodo, Farajallah, Perwitasari-Farajallah, and Hamidy, 2025, J. Trop. Biodiversity Biotechnol., 10 (jtbb13642): 1, formally removed the species from the synonymy of Nyctixalus pictus, on the basis of molecular phylogenetics and morphometrics. See comment under Nyctixalus pictus for additional relevant literature. Distinctiveness from Nyctixalus pictus rejected by Inger and Tan, 1996, Raffles Bull. Zool., 44: 564. Ibnudarda, Kadafi, Priambodo, Farajallah, Perwitasari-Farajallah, and Hamidy, 2025, J. Trop. Biodiversity Biotechnol., 10 (jtbb13642): 1, formally removed Nyctixalus anodon from the synonymy of Nyctixalus pictus, on the basis of molecular phylogenetics. Dever, McGuire, Iskandar, Hamidy, and Grismer, 2026, Zootaxa, 5777: 283–306, provided comparative morphology, morphometrics, and molecular markers (16S mtDNA) distinguishing the species.

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