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Nyctixalus margaritifer Boulenger, 1882
Nyctixalus margaritifer Boulenger, 1882, Ann. Mag. Nat. Hist., Ser. 5, 10: 35. Holotype: "Royal Museum, Brussels", now lost according to Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257, who designated as neotype BMNH 1885.12.31.35. Type locality: "purchased as being from the East Indies"; neotype from "Willis Mountains (Java)", Indonesia. Synonymy by Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257.
Ixalus flavosignatus Boettger, 1893, Zool. Anz., 16: 339. Holotype: SMF 7036 according to Mertens, 1967, Senckenb. Biol., 48(A): 47. Type locality: "Tjisurupan, West-Java", Indonesa. Synonymy (with Nyctixalus margaritifer) by Smith, 1931, Bull. Raffles Mus., 5: 17, and (with Ixalus pictus) by Inger, 1966, Fieldiana, Zool., 52: 349-350. Recognized (as Nyctixalus flavosignatus) without comment by Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257
Philautus flavosignatus — Smith, 1931, Zool. Rec., 67: 30. Mertens, 1967, Senckenb. Biol., 48(A): 47.
Philautus pictus margaritifer — Inger, 1966, Fieldiana, Zool., 52: 350.
Nyctixalus pictus margaritifer — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257; Bossuyt and Dubois, 2001, Zeylanica, 6: 31.
Nyctixalus flavosignatus — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257; Delorme, Dubois, Grosjean, and Ohler, 2005, Bull. Mens. Soc. Linn. Lyon, 74: 166. Recognized without discussion.
English Names
Java Indonesian Treefrog (Nycixalus flavosignatus [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 111).
Pearly Tree Frog (Iskandar, 1998, Amph. Java Bali: 84).
Distribution
Disjunct populationis on Java, Indonesia, in undisturbed highland forest above 700 m elevation.
Geographic Occurrence
Natural Resident: Indonesia
Endemic: Indonesia
Comment
Nyctixalus margaritifer considered possibly a synonym of Philautus aurifasciatus by Liem, 1973 "1971", Philipp. J. Sci., 100: 157. See discussion by Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 255-257, about confusion surrounding the application of the name Nyctixalus margaritifer. Iskandar, 1998, Amph. Java Bali: 84-85, provided a brief account and restriction of the name to the Javan population. See comments under Nyctixalus flavosignatus and Nyctixalus pictus. Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257, removed Ixalus flavosignatus from synonymy under Nyctixalus margaritifer; without discussion, implying that two species in this genus occur in Java. Iskandar, 1998, Amph. Java Bali: 84-85, did not recognize Nyctixalus flavosignatus as distinct from Nyctixalus margaritifer but did not address the issue. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 122-123, to compare this species to other rhacophorids of the Sunda Shelf region. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 516. See brief account for East Java by Amin, 2020, Frogs of East Java: 63–65.
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