Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nyctixalus margaritifer Boulenger, 1882

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Nyctixalus > Species: Nyctixalus margaritifer

Nyctixalus margaritifer Boulenger, 1882, Ann. Mag. Nat. Hist., Ser. 5, 10: 35. Holotype: "Royal Museum, Brussels", now lost according to Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257, who designated as neotype BMNH 1885.12.31.35. Type locality: "purchased as being from the East Indies"; neotype from "Willis Mountains (Java)", Indonesia. Synonymy by Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257.

Ixalus flavosignatus Boettger, 1893, Zool. Anz., 16: 339. Holotype: SMF 7036 according to Mertens, 1967, Senckenb. Biol., 48(A): 47. Type locality: "Tjisurupan, West-Java", Indonesa. Synonymy (with Nyctixalus margaritifer) by Smith, 1931, Bull. Raffles Mus., 5: 17, and (with Ixalus pictus) by Inger, 1966, Fieldiana, Zool., 52: 349-350. Recognized (as Nyctixalus flavosignatus) without comment by Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257

Philautus flavosignatus — Smith, 1931, Zool. Rec., 67: 30. Mertens, 1967, Senckenb. Biol., 48(A): 47.

Philautus pictus margaritifer — Inger, 1966, Fieldiana, Zool., 52: 350.

Nyctixalus pictus margaritifer — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257; Bossuyt and Dubois, 2001, Zeylanica, 6: 31.

Nyctixalus flavosignatus — Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257; Delorme, Dubois, Grosjean, and Ohler, 2005, Bull. Mens. Soc. Linn. Lyon, 74: 166. Recognized without discussion.

English Names

Java Indonesian Treefrog (Nycixalus flavosignatus [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 111).

Pearly Tree Frog (Iskandar, 1998, Amph. Java Bali: 84).

Distribution

Disjunct populations on Java, Indonesia, in undisturbed highland forest from 700 to 1200 m elevation.

Geographic Occurrence

Natural Resident: Indonesia

Endemic: Indonesia

Comment

Nyctixalus margaritifer considered possibly a synonym of Philautus aurifasciatus by Liem, 1973 "1971", Philipp. J. Sci., 100: 157. See discussion by Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 255-257, about confusion surrounding the application of the name Nyctixalus margaritifer. Iskandar, 1998, Amph. Java Bali: 84-85, provided a brief account and restriction of the name to the Javan population. See comments under Nyctixalus flavosignatus and Nyctixalus pictus. Dubois, 1981, Monit. Zool. Ital., N.S., Suppl., 15: 257, removed Ixalus flavosignatus from synonymy under Nyctixalus margaritifer; without discussion, implying that two species in this genus occur in Java. Iskandar, 1998, Amph. Java Bali: 84-85, did not recognize Nyctixalus flavosignatus as distinct from Nyctixalus margaritifer but did not address the issue. See identification table by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 122-123, to compare this species to other rhacophorids of the Sunda Shelf region. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 516. See brief account for East Java by Amin, 2020, Frogs of East Java: 63–65. Hanifa, Septiadi, Hasyim, Fathoni, Prabasari, Erfanda, Rahmadana, Widjanarko, and Kurniawan, 2023, Check List, 19: 971–982, provided new records from central and eastern Java and also discussed morphological variation. Ibnudarda, Farajallah, Perwitasari, Hamidy, Prihatini, and Darmawan, 2024, Zootaxa, 5410: 297–300, reported on the advertisement call and reciprocal calling of the sexes.

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For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.