Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Telmatobius verrucosus Werner, 1899

Class: Amphibia > Order: Anura > Family: Telmatobiidae > Genus: Telmatobius > Species: Telmatobius verrucosus

Telmatobius verrucosus Werner, 1899, Zool. Anz., 22: 482. Holotype: NHMW 22921, acording to De la Riva, 2005, in Lavilla and De la Riva (eds.), Monogr. Herpetol., 7: 88, who noted that the previous statements about NHMW 22922 being the type (e.g., Häupl and Tiedemann, 1978, Kat. Wiss. Samml. Naturhist. Mus. Wien, 2: 29, and Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 34) are in error. Type locality: "Chaco, Bolivia"; corrected to Chaco, on the Unduavi River, 25 km northwest of Puente de la Via, La Paz, Bolivia, by Schmidt, 1954, Fieldiana, Zool., 34: 284. De la Riva, 2005, in Lavilla and De la Riva (eds.), Monogr. Herpetol., 7: 88, suggested that the actual type locality must be at higher elevation than the stated location.

Telmatobius marmoratus verrucosus — Vellard, 1969 "1968", Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 40: 1111.

Telmatobius jahuira Lavilla and Ergueta-Sandoval, 1995, Alytes, 13: 46. Holotype: CBF 01675, by original designation. Type locality: "el Río Chairo, en las proximadades de la Mina Copacabana (aproximadamente 16° 16′ S 67° 50′ W), Kkota Pata, Departamento La Paz, Bolivia". Synonymy by De la Riva, 2005, IN Lavilla and De la Riva (eds.), Monogr. Herpetol., 7: 88.

Telmatobius verrucosus — De la Riva, Köhler, Lötters, and Reichle, 2000, Rev. Esp. Herpetol., 14: 50. De la Riva, 2005, IN Lavilla and De la Riva (eds.), Monogr. Herpetol., 7: 86.

Common Names

Chaco Water Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 85).

Distribution

Hillsides of Mount Illimani (Lambate) to the Quelhuacota Valley, near Charazani, a range of about 2000 km, in the Department of La Paz, Bolivia, 2900-3800 m elevation.

Geographic Occurrence

Natural Resident: Bolivia

Endemic: Bolivia

Comment

Removed from the synonymy of Telmatobius marmoratus by De la Riva, Köhler, Lötters, and Reichle, 2000, Rev. Esp. Herpetol., 14: 50, and De la Riva, 2005, in Lavilla and De la Riva (eds.), Monogr. Herpetol., 7: 86-89 (who provided an account), where it had been placed by Vellard, 1969 "1968", Bull. Mus. Natl. Hist. Nat. Paris, Ser. 2, 40: 1112. De la Riva, 2005, in Lavilla and De la Riva (eds.), Monogr. Herpetol., 7: 88-89, discussed errors in the literature due to the misidentification of the type and other serious problems in the history of delimiting this taxon. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 419.

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For additional information see AmphibiaWeb report
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For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.