Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Isthmura Dubois and Raffaelli, 2012

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Hemidactyliinae > Genus: Isthmura

7 species

Isthmura Dubois and Raffaëlli, 2012, Alytes, 28: 77–161. Type species: Spelerpes belli Gray, 1850, by original designation. Coined as a subgenus of Pseudoeurycea in the sense of excluding Ixalotriton.

Isthmura — Rovito, Vázquez-Almazán, Papenfuss, Parra-Olea, and Wake, 2015, Zool. J. Linn. Soc., 175: 185, treatment as a genus.

Distribution

Montane Mexico from east-central Sonora and adjacent Chihuahua in the Sierra Madre Occidental and from southern Tamaulipas in the Sierra Madre Oriental south through Nayarite, and Zacatecas,to Tlaxcala, Hidalgo, wester-central Veracruz, and the Sierra Madre del Sur of Guerrero and Oaxaca.

Comment

Formerly the Pseudoeurycea bellii group, removed from that genus by Rovito, Parra-Olea, Recuero, and Wake, 2015, Zool. J. Linn. Soc., 175: 184, to preserve the monophyly of Pseudoeurycea. Bryson, Zarza, Grummer, Parra-Olea, Flores-Villela, Klicka, and McCormack, 2018, Mol. Phylogenet. Evol., 125: 75–84, provided a tree and discussed the phylogenomic diversification across the Mexican highlands.

Contained taxa (7 sp.):

Isthmura bellii (Gray, 1850)

Isthmura boneti (Alvarez and Martín, 1967)

Isthmura corrugata Sandoval-Comte, Pineda-Arredondo, Rovito, and Luría-Manzano, 2017

Isthmura gigantea (Taylor, 1939)

Isthmura maxima (Parra-Olea, García-París, Papenfuss, and Wake, 2005)

Isthmura naucampatepetl (Parra-Olea, Papenfuss, and Wake, 2001)

Isthmura sierraoccidentalis (Lowe, Jones, and Wright, 1968)

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