Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Desmognathus folkertsi Camp, Tilley, Austin, and Marshall, 2002

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Plethodontinae > Genus: Desmognathus > Species: Desmognathus folkertsi

Desmognathus folkertsi Camp, Tilley, Austin, and Marshall, 2002, Herpetologica, 58: 477. Holotype: USNM 536397, by original designation. Type locality: "south of Wolf Creek Road (34° 46′ 05″ N latitude; 83° 56′ 37″ W longitude) on an upper tributary of the West Fork of Wolf Creek at an elevation of 834 m, Union County, Georgia", USA.

Desmognathus (Leurognathus) folkertsi — Dubois and Raffaëlli, 2012, Alytes, 28: 145. See comment under Desmognathus regarding the status of this subgenus.

Common Names

Dwarf Black-bellied Salamander (Camp, Marshall, Landau, Austin, and Tilley, 2000, Copeia, 2000: 477; Tilley, Highton, and Wake, 2012, in Crother (ed.), Herpetol. Circ., 39: 16; Tilley, Highton, and Wake, 2012, in Crother (ed.), Herpetol. Circ., 39: 25; Powell, Conant, and Collins, 2016, Field Guide Rept. Amph. E. North Am., 4th ed.: 44).

Dwarf Black-belly Salamander (Camp, 2004, Cat. Am. Amph. Rept., 782: 1).

Dwarf Blackbelly Salamander (Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 11).

Dwarf Black-bellied Dwarf Salamander (Raffaëlli, 2022, Salamanders & Newts of the World: 1011).

Distribution

Tributaries of the West Fork of Wolf Creek and Helton Creek, both in Union County, Georgia, as well in adjacent Clay County, North Carolina, USA.

Geographic Occurrence

Natural Resident: United States of America, United States of America - Georgia, United States of America - North Carolina

Endemic: United States of America

Comment

Most similar to Desmognathus quadramaculatus (now composed of Desmognathus amphileucus, Desmognathus gvnigeusgwotli, Desmognathus mavrokoilius, and Desmognathus kanawha) according to the original publication. See account by Camp, 2004, Cat. Am. Amph. Rept., 782: 1–3. First record for North Carolina by Rothermel, Jensen, Camp, and Schwaner, 2007, Herpetol. Rev., 38: 213. Wooten, Camp, and Rissler, 2010, Conserv. Genetics, 11: 835–854, reported on genetic variation across the range of the species. Camp, 2005, in Lannoo (ed.), Amph. Declines: 706–708, provided an account containing a detailed summary of the literature and range. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 431, provided a brief account, photograph, and range map. Beamer and Lamb, 2020, Zootaxa, 4734: 1–61, in their discussion of Desmognathus mtDNA phylogenetics, noted this species to sit phylogenetically within the Desmognathus marmoratus/Desmognathus 'quadramaculatus' complex of cryptic species. Pyron, O'Connell, Lemmon, Lemmon, and Beamer, 2022, Ecol. Evol., 12 (2: e8574): 1–38, provided molecular evidence that Desmognathus folkertsi is genealogically cohesive. Raffaëlli, 2022, Salamanders & Newts of the World: 1011, provided an account summarizing systematics, morphology, life history, population status, and distribution (including a polygon map). Camp, Felix, and Wooten, 2022, Amphibia-Reptilia, 43: 133–140, reported on morphological homoplasy among semi-aquatic species (Desmognathus welteri, Desmognathus folkertsi, northern "quadramaculatus" (now Desmgnathus kanawha), and southern "quadramaculatus" (now Desmognathus amphileucus).

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.