Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Desmognathus intermedius (Pope, 1928)

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Plethodontinae > Genus: Desmognathus > Species: Desmognathus intermedius

Leurognathus marmorata intermedia Pope, 1928, Am. Mus. Novit., 306: 14. Holotype: AMNH 25557, by original designation. Type locality: "Davis Gap, Waynesville, North Carolina", USA; emended by Bishop, 1943, Handb. Salamanders: 224, to "Davis Farm, about 2 miles east and a little north of Waynesville, (Haywood County) on Highway No. 276." Status rejected by Martof, 1962, Am. Midl. Nat., 67: 30.

Leurognathus intermedia — Pope and Hairston, 1947, Fieldiana, Zool., 31: 160.

Leurognathus marmorata melania Martof, 1956, Occas. Pap. Mus. Zool. Univ. Michigan, 575: 6. Holotype: UMMZ 111564, by original designation. Type locality: "Otter Creek, tributary of the Nantahala River, 0.5 of a mile west of Tellico Gap, Macon County, North Carolina; 3,600 feet elevation", USA. Status rejected by Martof, 1962, Am. Midl. Nat., 67: 30. Synonymy by Raffaëlli, 2022, Salamanders & Newts of the World: 1013.

Leurognathus marmoratus melanius — Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 229.

Desmognathus (Leurognathus) melanius — Dubois and Raffaëlli, 2012, Alytes, 28: 145.

Desmognathus (Leurognathus) intermedius — Raffaëlli, 2022, Salamanders & Newts of the World: 1013.

English Names

Black Shovel-nosed Salamander (Conant, 1958, Field Guide Rept. Amph. E. Cent. N. Am.: 229).

Otter Creek Dusky Salamander (Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 178).

Central Shovel-nosed Dusky Salamander (Raffaëlli, 2022, Salamanders & Newts of the World: 1013).

Southern Shovel-nosed Salamander (Schmidt, 1953, Check List N. Am. Amph. Rept., Ed. 6: 32; Conant, Cagle, Goin, Lowe, Neill, Netting, Schmidt, Shaw, Stebbins, and Bogert, 1956, Copeia, 1956: 174).

Pope's Salamander (Bishop, 1943, Handb. Salamanders: 224).

Distribution

Nantahala, Cheoh, Oconaluftee, Little Pigeon, Pigeon, and Tuckasegee River drainages of Appalachian western North Carolina and adjacent Tennessee, USA.

Comment

See comments under Desmognathus marmoratus, where literature for when Desmognathus intermedius was considered part of that taxon. Desmognathus intermedius is the Desmognathus quadramaculatus C of Beamer and Lamb, 2020, Zootaxa, 4734: 1–61, as discussed by Raffaëlli, 2022, Salamanders & Newts of the World: 1013–1014, who provided a detailed account of its range, natural history, and literature.

External links:

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For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.