Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Plethodon websteri Highton, 1979

Class: Amphibia > Order: Caudata > Family: Plethodontidae > Subfamily: Plethodontinae > Genus: Plethodon > Species: Plethodon websteri

Plethodon websteri Highton, 1979, Brimleyana, 1: 32. Holotype: USNM 204814, by original designation. Type locality: "0.6 km east, 0.9 km south of Howelton, Etowah County, Alabama", USA.

Plethodon (Plethodon) websteri — Vieites, Nieto-Roman, Wake, and Wake, 2011, Mol. Phylogenet. Evol., 59: 632, by implication.

English Names

Webster's Salamander (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 33; Collins, 1997, Herpetol. Circ., 25: 9; Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 29; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 14; Tilley, Highton, and Wake, 2012, in Crother (ed.), Herpetol. Circ., 39: 30; Highton, Bonett, and Jockusch, 2017, in Crother (ed.), Herpetol. Circ., 43: 32).

Distribution

Known from central Alabama and west-central Georgia; apparently disjunct populations occur in Clarke County (Alabama), Hinds and Winston counties (Mississippi), West Feliciana Parish (Louisiana), and McCormick County (South Carolina), USA.

Comment

In the Plethodon welleri group of Highton and Larson, 1979, Syst. Zool., 28: 579–599. See accounts by Highton, 1986, Cat. Am. Amph. Rept., 384: 1–2, and Petranka, 1998, Salamand. U.S. Canada: 407–409. Forms a narrow hybrid zone with Plethodon dorsalis (Highton, 1985, J. Herpetol., 19: 544–546). Beamer and Lannoo, 2005, in Lannoo (ed.), Amph. Declines: 852–853, provided a detailed account that summarized the biology and conservation literature. Highton, Hastings, Palmer, Watts, Hass, Culver, and Arnold, 2012, Mol. Phylogenet. Evol., 63: 278–290, noted 12S mtDNA representative of the Plethodon glutinosus complex in Plethodon websteri, but that other genes and allozymes placed Plethodon websteri outside of the Plethodon glutinosus complex. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 393, provided a brief account, photograph, and range map. Feist, Mann, Graham, Wooten, Toyota, Mann, Balius, Polanco, Wolwehender, and Moore, 2019, Conserv. Genetics, 20: 947–960, suggested on the basis of mtDNA analysis that the nominal species may be composed of two lineage-species and that it is genetically distant from Plethodon welleri and not part of that group.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.