Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Proteus bosniensis Šunje, Jelič, Zimič, Dufresnes and Jablonski, 2026

Class: Amphibia > Order: Caudata > Family: Proteidae > Genus: Proteus > Species: Proteus bosniensis

Proteus bosniensis Šunje, Jelič, Zimič, Dufresnes and Jablonski in Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 39. Holotype NMBH 11568, by original designation. Type locality: "Suvaja, Lušci Palanka, Bosnia and Herzegovina". ZooBank Publication registration: 9EE06D32-44D5-48DB-A34E-A9BB7849433D (13 August 2026)

Proteus bosniacus Trontelj, Gorički, Konstanjšek, Zakšek, Delić, Bressi, Mauri, Lukić Bilela, Jalžić, and Recknagel, 2026, Nat. Sloveniae, 28: 21. Holotype: ULBF PA170, by original designation. Type locality: "Cave Suvaja Pećina, about 6 km SE of Lušci Palanka, Sanski Most, Bosnia and Herzegovina; Lat., Lon. = 44.7006, 16.4713". ZooBank Publication registration: D60B4EF9-D0CF-46C3-B015-467D6975F0E1 (24 August 2026).

Common Names

Bosnian Olm (Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 39).

Bosanska čovječja ribica (Bosnian: Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 39).

Distribution

Restricted to the upper reaches of the Una River and Sana River drainages within the Danube Basin of northwestern Bosnia and Herzegovina, primarily surrounding Grmeč; the species inhabits a complex karst landscape composed of a series of high karst poljes (approximately 600 m elevation) that drain through extensive underground systems into lower poljes (around 300 m elevation), emerging through large karst springs such as Klokot, Krušnica, Sanica and Dabar.

Geographic Occurrence

Natural Resident: Bosnia and Herzegovina

Endemic: Bosnia and Herzegovina

Comment

The Bosanska Krajina clade of Trontelj, Douady, Fišer, Gilbert, Gorički, Lefébure, Sket, and Zakšek, 2009, Freshwater Biol., 54: 737 and the Krajina lineage of Kostanjšek, Zakšek, Bizjak-Mali, and Trontelj, 2023, In Mallard et al. (eds.), Groundwater Ecology and Evolution. 2nd ed: 310, and of the revisionary works of Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 1–51, and Trontelj, Gorički, Konstanjšek, Zakšek, Delić, Bressi, Mauri, Lukić Bilela, Jalžić, and Recknagel, 2026, Nat. Sloveniae, 28: 11–27, where comparative morphology, morphometrics, molecular markers (mt- and nuDNA), a dot map of the distribution, habitat, and conservation status were provided.

External links:

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For related information on conservation and images as well as observations see iNaturalist