Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Proteus carrarae (Fitzinger, 1850)

Class: Amphibia > Order: Caudata > Family: Proteidae > Genus: Proteus > Species: Proteus carrarae

Hypochthon Carrarae Fitzinger, 1850, Sitzungsber. Akad. Wiss. Wien, Phys. Math. Naturwiss. Kl., 5: 300. Syntypes: Not stated, although clearly NHMW; including NHMW 19980, according to Häupl, Tiedemann, and Grillitsch, 1994, Kat. Wiss. Samml. Naturhist. Mus. Wien, 9: 13, and Gemel, Gassner, and Schweiger, 2019, Ann. Naturhist. Mus. Wien, Ser. B, 121: 40. NHMW 19980/1 designated lectotype by Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 25. Type locality]ies: "Sign [= Sinj ] und der Narenta", Slovenia. Restricted to Sinj by lectotype designation

Proteus carrarae — Cope, 1866, J. Acad. Nat. Sci. Philadelphia, Ser. 2, 6: 103; Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 33; Trontelj, Gorički, Konstanjšek, Zakšek, Delić, Bressi, Mauri, Lukić Bilela, Jalžić, and Recknagel, 2026, Nat. Sloveniae, 28: 12.

Proteus anguinus var. carrarae — Boulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 86.

Common Names

Dalmatian Olm (Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 32).

Dalmatinska čovječja ribica (Croatian: Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 32).

Hercegovačka čovječja ribica (Bosnian: Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 32).

Distribution

Extends along the coastal Dinaric Karst from northern Dalmatia to southern Herzegovina and the Dubrovnik hinterland possibly extending as far south as the Bay of Kotor if an uncertain eDNA record from Sopot (Risan), Montenegro, is confirmed.

Geographic Occurrence

Natural Resident: Bosnia and Herzegovina, Croatia

Likely/Controversially Present: Montenegro

Comment

The Para-littoral lineage, the sister of Proteus bosniensis of the original publication, where comparative morphology, morphometrics, molecular markers (mt- and nuDNA), a dot map of the distribution, habitat, and conservation status were provided. Removed from the synonymy of Proteus anguinus by Dufresnes, Monod-Broca, Jelić, Šunje, Manenti, Barzaghi, Zimić, Chouchane, Denoël, Ohler, and Jablonski, 2026, Amphibia-Reptilia, Published online ahead of print: 32, where it had been placed by Gray, 1850, Cat. Spec. Amph. Coll. Brit. Mus., Batr. Grad.: 65, and Boulenger, 1882, Cat. Batr. Grad. Batr. Apoda Coll. Brit. Mus., Ed. 2: 85. Validity had been doubted by implication of Schreiber, 1875, Herpetol. Eur.: 13–14. Gorički, Stanković, Snoj, Kuntner, Jeffery, Trontelj, Pavićević, Grizelj, Năpăruş-Aljančič, and Aljančič, 2017, Sci. Rep. (Nature, London), 7 (45054): 1–11, provided an environmental DNA record from Montenegro that requires confirmation.

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