Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Lissotriton graecus (Wolterstorff, 1906)

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Lissotriton > Species: Lissotriton graecus

Triton vulgaris graeca Wolterstorff, 1906, Zool. Anz., 29: 137. Syntypes: MM; destroyed in W.W.II. SMF 1038 (presumably exchanged from MM) designated lectotype by Mertens, 1967, Senckenb. Biol., 48(A): 38. Type locality: "Korfu", Greece.

Triton vulgaris intermedia Kolombatovic, 1907, Glasn. Hrvatsk. Naravosl. Drustva, Zagreb, 19: 13. Holotype: Not stated or known to exist. Type locality: "Karakasika al nord di Sinj", Dalmatia, Croatia. Placed in this synonymy provisionally on the basis of geography (DRF).

Triton vulgaris graeca forma tomasinii Wolterstorff, 1908, Lacerta, 6: 23. Syntypes: MM, by original designation; destroyed in W.W.II according to Brame, 1972, Checklist Living & Fossil Salamand. World (Unpubl. MS): 109. Type locality: "Teodo di Cattaro", Dalmatia, Croatia. Unavailable tetranominal.

Triton vulgaris graeca forma corcyrensis Wolterstorff, 1908, Lacerta, 6: 23. Syntypes: MM, by original designation; destroyed in W.W.II according to Brame, 1972, Checklist Living & Fossil Salamand. World (Unpubl. MS): 109. Type locality: "Korfu" Island, Greece. Unavailable tetranominal.

Triton meridionalis var. graeca — Schreiber, 1912, Herpetol. Eur., Ed. 2: 82.

Triton vulgaris graeca — Wolterstorff, 1912, Bl. Aquar. Terrarienkd., Stuttgart, 23: 190.

Triton meridionalis var. graeca forma tomasinii — Schreiber, 1912, Herpetol. Eur., Ed. 2: 79.

Triton vulgaris forma schreiberi Wolterstorff, 1914, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 2: 371. Syntypes: MM (all except lectotype destroyed in WWII); MM 615 designated lectotype by Bischoff, 1977, Bonn. Zool. Beitr., 28: 117-121. Type locality: "Bokanjacko blato, einem grossen Sumpf bei Zara", Croatia. Placed in this synonymy provisionally on the basis of geography (DRF).

Triton vulgaris forma schreiberi — Wolterstorff, 1925, Abh. Ber. Mus. Nat. Heimatkd. Magdeburg, 4: 275.

Triton (Palaeotriton) vulgaris graecus — Bolkay, 1927, Glasn. Zemaljskog Muz. Bosni Hercegov., 39: 63. Bolkay, 1928, Z. Anat. Entwicklungesch., 86: 259.

Triturus vulgaris graecus — Mertens and Müller, 1928, Abh. Senckenb. Naturforsch. Ges., 41: 13.

Triturus vulgaris tomasinii — Mertens and Müller, 1928, Abh. Senckenb. Naturforsch. Ges., 41: 13. Mertens and Müller, 1940, Abh. Senckenb. Naturforsch. Ges., 451: 11.

Triturus vulgaris schreiberi — Mertens and Müller, 1940, Abh. Senckenb. Naturforsch. Ges., 451: 11.

Triturus meridionalis graeca — Bodenheimer, 1944, Istanbul Univ. Fen. Fak. Mec., Seri B, 9: 6.

Lissotriton graecus — Dubois and Raffaëlli, 2009, Alytes, 26: 40; Pabijan, Zieliński, Dudek, Stuglik, and Babik, 2017, Mol. Phylogenet. Evol., 116: 1; Wielstra, Canestrelli, Cvijanović, Denoël, Fijarczyk, Jablonski, Liana, Naumov, Olgun, Pabijan, Pezzarossa, Popgeorgiev, Salvi, Si, Sillero, Sotiropoulos, Zieliński, and Babik, 2018, Amphibia-Reptilia, 39: 252.

Lissotriton (Lissotriton) (vulgaris) graecus — Dubois and Raffaëlli, 2009, Alytes, 26: 66.

English Names

Greek Newt (Steward, 1969, Tailed Amph. Eur.: 40).

Schreiber's Smooth Newt (Steward, 1969, Tailed Amph. Eur.: 40).

Greek Smooth Newt (Wielstra, Canestrelli, Cvijanović, Denoël, Fijarczyk, Jablonski, Liana, Naumov, Olgun, Pabijan, Pezzarossa, Popgeorgiev, Salvi, Si, Sillero, Sotiropoulos, Zieliński, and Babik, 2018, Amphibia-Reptilia, 39: 254).

Distribution

Dalmatian coast of central Croatia, south through Bosnia and Herzegovina, Montenegro, southern Kosovo, North Macedonia, and Albania to and including most of Greece.

Comment

Razzetti, Andreone, Corti, and Sindaco, 2006, in Sindaco et al. (ed.), Atlas Ital. Amph. Rept.: 162, suggested that evidence provided by Pellarini and Lapini, 2000, Atti del I Congr. Naz. Soc. Herpetol. Italica, Torino, 1996, Atti del I Congr. Naz. della Soc. Herpetol. Italica, Mus. Reg. Sci. Nat. Torino: 347–351, suggested that Lissotriton vulgaris vulgaris and Lissotriton vulgaris meridionalis may be distinct species. This was formalized by Dubois and Raffaëlli, 2009, Alytes, 26: 39–40. Valakos, Pafilis, Sotiropoulos, Lymberakis, Maragou, and Foufopoulos, 2008, Amph. Rept. Greece: 151–154, provided an account for Greece (as Lissotriton vulgaris). Documented zone of intergradation/hybridization on the eastern part of its range with Lissotriton vulgaris; see comment under Lissotriton vulgaris. Speybroeck, Beukema, and Crochet, 2010, Zootaxa, 2492: 3, discussed the problematic nature of the evidence supporting recognition of this taxon and refrained from recognizing it as a species distinct from Lissotriton vulgaris in part based on an adherence to the biological species concept. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 122–123, provided a brief account, photo, and map. Wielstra, Canestrelli, Cvijanović, Denoël, Fijarczyk, Jablonski, Liana, Naumov, Olgun, Pabijan, Pezzarossa, Popgeorgiev, Salvi, Si, Sillero, Sotiropoulos, Zieliński, and Babik, 2018, Amphibia-Reptilia, 39: 252–259, reviewed the relevant taxonomic literature of the Lissotriton vulgaris complex and provided a range map, noting the narrow hybrid zone with Lissotriton vulgaris. See Dufresnes, 2019, Amph. Eur., N. Afr., & Middle East: 123, for brief summary of identifying morphology and biology, a range map, as well as a photograph. See Speybroeck, Beukema, Dufresnes, Fritz, Jablonski, Lymberakis, Martínez-Solano, Razzetti, Vamberger, Vences, Vörös, and Crochet, 2020, Amphibia-Reptilia, 41: 144, for taxonomic comments.

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