Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Tylototriton hainanensis Fei, Ye, and Yang, 1984

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Pleurodelinae > Genus: Tylototriton > Species: Tylototriton hainanensis

Tylototriton hainanensis Fei, Ye, and Yang, 1984, Acta Zool. Sinica, 30: 85-89. Holotype: CIB 64III1379, by original designation. Type locality: "Nalong, Wuzhi Shan, alt. 770 m", Hainan, China.

Pleurodeles (Tylototrion) hainanensis — Risch, 1985, J. Bengal Nat. Hist. Soc., N.S.,, 4: 142.

Pleurodeles (Echinotriton) hainanensis — Dubois, 1987 "1986", Alytes, 5: 11.

Tylototriton (Yaotriton) hainanensis — Dubois and Raffaëlli, 2009, Alytes, 26: 68.

Yaotriton hainanensis — Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 91; Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 408.

English Names

Hainan Knobby Newt (Fei, 1999, Atlas Amph. China: 40).

Distribution

Biasha, Leong, Lingshui and Qiongzhong, Hainan Island, China, 770–970 m elevation. See comment.

Comment

See accounts by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 74, and Fei, 1999, Atlas Amph. China: 40. Synonymy with Tylototrion asperrimus by Zhao, Hu, Jiang, and Yang, 1988, Studies on Chinese Salamanders: 63; this rejected by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China 74. In the Tylototriton asperrimus group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 42 (although they only addressed Chinese species). See comments under Tylototriton and Tylototriton asperrimus. Fei, Hu, Ye, and Huang, 2006, Fauna Sinica, Amph. 1: 261–265, provided an account. See photograph, map, description of geographic range and habitat, and conservation status in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 607. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 76, provided a brief account including photographs. Shi, 2011, Amph. Rept. Fauna Hainan: 36–38, provided an account. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 91, provided an account, photographs, and a map. Raffaëlli, 2013, Urodeles du Monde, 2nd ed.: 186, provided a brief account, photograph, and map. Sparreboom, 2014, Salamanders Old World: 361–362, reviewed the biology, characteristics, distribution, reproduction, and conservation of the species. Fei and Ye, 2016, Amph. China, 1: 301–303, provided an account (as Yaotriton hainanensis), photographs, and range map. Bernardes, Le, Nguyen, Pham, Pham, Nguyen, Rödder, Bonkowski, and Ziegler, 2020, ZooKeys, 935: 121–164, provided a range map and mapped a population in western Guizhou, China, as Tylototriton cf. hainanensis, implying an unnamed species. In the Tylototriton (Yaotriton) asperrimus species group of Poyarkov, Nguyen, and Arkhipov, 2021, Taprobanica, 10: 4–22, who discussed phylogenetics.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.