Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Mertensiella djanaschvilii Tartarashvili and Bakradze, 1989

Class: Amphibia > Order: Caudata > Family: Salamandridae > Subfamily: Salamandrinae > Genus: Mertensiella > Species: Mertensiella djanaschvilii

Mertensiella caucasica djanaschvilii Tartarashvili and Bakradze, 1989, Bull. Acad. Sci. Georgia, Tbilisi, 133: 177-179. Types: Not stated. Type locality: "in Georgian SSR in the vicinity of mount Tirala (Batumi region)", Georgia. Stated as "Mtirala Mountain, environs of Batumi Town, Adjaria, Georgia" by Tuniyev, 1999, Adv. Amph. Res. Former Soviet Union, 3: 48.

Mertensiella djanaschvilii — Raffaëlli, 2022, Salamanders & Newts of the World: 255.

English Names

None noted.

Distribution

Western Caucasus from the Borjomi area, western Georgia, to northeastern Turkey to the west in the eastern Black Sea region, west to Çambaşi, south of Ordu (Ordu, Gresun, Rize, Trabzon, Artvin, Kars, Bayburt, Gümüshane provinces).

Geographic Occurrence

Natural Resident: Georgia, Turkey

Comment

Başoğlu and Özeti, 1973, Türkiye Amphibileri: 36–37, provided an account for the Turkish populations (as Mertensiella caucasica). Özeti and Yilmaz, 1994, Türkiye Amfibileri: 51–54, provided an account for Turkish populations (as Mertensiella caucasica). Tuniyev, 1999, Adv. Amph. Res. Former Soviet Union, 3: 48-49, discussed the status and distribution of Mertensiella caucasica djanaschvilii. Beşir and Gül, 2019, Herpetol. Notes, 12: 45–51, provided a dot map for Rize Province, northeastern Turkey (as Mertensiella djanaschvilii). See comments under Mertensiella caucasica noting literature that included this taxon as a subspecies of that species. Kurnaz and Kutrup, 2019, Nat. Conserv. Res., 4: 109–111, provided new localities and modeled the distribution in Gümüşhane province in Turkey (as Mertensiella caucasica). Çiçek, Yakin, Afsar, Ayaz, and Tok, 2019, Acta Biol. Turcica, 32: 37–41, provided new localities in Turkey and a dot map for the species' range (as Mertensiella caucasica). Yaşar, Çiçek, Mulder, and Tok, 2021, North-West. J. Zool., Romania, 17(e201512): 232–275, discussed and grid-mapped the range in Turkey (as Mertensiella caucasica). Raffaëlli, 2022, Salamanders & Newts of the World: 255, resurrected the name Mertensiella djanaschvilii on the basis of evidence presented by Tarkhnishvili, Thorpe, and Arntzen, 2000, Mol. Phylogenet. Evol., 14: 414–422, and Tarkhnishvili, Kaya, Gavashelishvili, and Serbinova, 2008, Herpetol. J., 18: 155–163.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.