Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Leptopelis rugosus (Ahl, 1924)

Class: Amphibia > Order: Anura > Family: Arthroleptidae > Subfamily: Leptopelinae > Genus: Leptopelis > Species: Leptopelis rugosus

Pseudocassina rugosa Ahl, 1924, Mitt. Zool. Mus. Berlin, 11: 9. Holotype: ZMB registration not provided, according to the original publication; ZMB 26915, according to Goutte, Reyes-Velasco, Kassie Teme, and Boissinot, 2022, ZooKeys, 1128: 76. Type locality: "Meta, Meta . . . Kolla, Zindelevat", Ethiopia. Given as "Meta, Kolla (approx. 9.17°, 38.25°, 2650 m a.s.l.)" by Goutte, Reyes-Velasco, Kassie Teme, and Boissinot, 2022, ZooKeys, 1128: 76.

Pseudocassina ocellata Ahl, 1924, Mitt. Zool. Mus. Berlin, 11: 8. Syntypes: ZMB (2 specimens), registration numbers not provided in the original publication' ZMB 26913 designated lectotype by Goutte, Reyes-Velasco, Kassie Teme, and Boissinot, 2022, ZooKeys, 1128: 76. Type localities: "Hochebene Didda" and "Somaliland". Actually in Ethiopia according to Tiutenko and Zinenko, 2021, ZooKeys, 1023: 143. Junior secondary homonym of Hylambates ocellatus Mocquard, 1902 (= Leptopelis ocellata), when both are in Leptopelis. Synonymy by Goutte, Reyes-Velasco, Kassie Teme, and Boissinot, 2022, ZooKeys, 1128: 76.

Leptopelis rugosus — Ahl, 1931, Das Tierreich, 55: 222; Goutte, Reyes-Velasco, Kassie Teme, and Boissinot, 2022, ZooKeys, 1128: 76.

Leptopelis montanus Tiutenko and Zinenko, 2021, ZooKeys, 1023: 143. Replacement name for Pseudocassina ocellatus Ahl, 1924.

English Names

Ocellated Burrowing Treefrog (Leptopelis montanus [no longer recognized]: Tiutenko and Zinenko, 2021, ZooKeys, 1023: 143).

Distribution

Grassy meadows of the Ethiopian Highlands north of the Great Rift Valley at mid- to high elevations (2339–3337 m) from near Debre Birhan, Debre Sina, Fiche, Addis Ababa, Holeta and Ambo; northernmost population was found near Mehal Meda, while the southernmost individuals were found between Ambo and Wonchi; Arussi Plateau record, east of the Great Rift Valley requires confirmation.

Geographic Occurrence

Natural Resident: Ethiopia

Endemic: Ethiopia

Comment

Removed from the synonymy of Leptopelis gramineus, by Tiutenko and Zinenko, 2021, ZooKeys, 1023: 143, who provided information on external morphology, genetic markers, and advertisement call as well as mapping the range, where it had been placed by Largen, 1977, Monit. Zool. Ital., N.S., Suppl., 9: 86. Earlier synonymy with Leptopelis rugosus by Ahl, 1931, Das Tierreich, 55: 222. Pseudocassina rugosa Ahl was placed in the synonymy of Leptopelis gramineus by Largen, 1977, Monit. Zool. Ital., N.S., Suppl., 9: 86; resurrected by Goutte, Reyes-Velasco, Kassie Teme, and Boissinot, 2022, ZooKeys, 1128: 76, who recognized it as an older name for Leptopelis montanus. These authors also summarized the relevant literature, mtDNA relationships, and morphometrics.

External links:

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For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.