Bombina fortinuptialis Hu and Wu, 1978

Class: Amphibia > Order: Anura > Family: Bombinatoridae > Genus: Bombina > Species: Bombina fortinuptialis

Bombina fortinuptialis Hu and Wu In Liu, Hu, Tian, and Wu, 1978, Mater. Herpetol. Res., 4: 18. Holotype: CIB 601750, by original designation. Type locality: Yangliuchong, Jinxiu, Yaoshan, Guangxi, altitude 1350 m, China. 

Bombina fortinuptialis Tian and Wu In Hu, Tian, and Wu, 1981, Acta Herpetol. Sinica, 5 (17): 111. Holotype: CIB 601750, by original designation. Type locality: Yangliuchong, Jinxiu, Yaoshan, Guangxi, altitude 1350 m, China. Unnecessary second description. 

Bombina (Glandula) fortinuptialis — Tian and Hu, 1985, Acta Herpetol. Sinica, Chengdu, N.S.,, 4 (3): 220.

Bombina (Grobina) fortinuptialis — Dubois, 1987 "1986", Alytes, 5: 97.

English Names

Guangxi Firebelly Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 38).

Large-spined Bell Toad (Fei, 1999, Atlas Amph. China: 58).


Eastern Guangxi Province (Jinxiu, Longsheng, and Pingnan), China.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of


See redescription by Tian and Wu in Hu, Tian, and Wu, 1981, Acta Herpetol. Sinica, 5 (17): 111–120. Zhao and Adler, 1993, Herpetol. China: 115, discussed authorship. See  brief account, map, and figure by Fei, 1999, Atlas Amph. China: 58–59. Zhang and Wen, 2000, Amph. Guangxi: 38, provided an account for Guangxi. Considered an allopatric population of Bombina maxima by Yu, Yang, Zhang, and Rao, 2007, J. Herpetol., 41: 365–377. See photograph, map, description of geographic range and habitats, and conservation status by Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 155. Zheng, Fu, and Li, 2009, Mol. Phylogenet. Evol., 52: 70–83, recognized Bombina microdeladigitoraBombina lichuanensis, and Bombina fortinuptialis as separate species and provided molecular evidence consistent with this view. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 106–108, provided a brief account. Pabijan, Wandycz, Hofman, Węcek, Piwczyński, and Szymura, 2013, Mol. Phylogenet. Evol., 69: 63–74, questioned the distinctivness of Bombina microdeladigitora, Bombina lichuanensis, and Bombina fortinuptialis due to low mtDNA distances between those populations. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 126–134, provided an account, photograph, and map.

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