Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rulyrana mcdiarmidi (Cisneros-Heredia, Venegas, Rada, and Schulte, 2008)

Class: Amphibia > Order: Anura > Family: Centrolenidae > Subfamily: Centroleninae > Genus: Rulyrana > Species: Rulyrana mcdiarmidi

Cochranella mcdiarmidi Cisneros-Heredia, Venegas, Rada, and Schulte, 2008, Herpetologica, 64: 342. Holotype: DFCH-USFQ D132, by original designation. Type locality: "along a small rivulet tributary of the Jambue River, ca. 6 km S from Zamora (ca. 04° 03′S, 78° 56′W, 1150 m), on the western slope of Contrafuerte de Tzunantza, Cordillera Oriental, eastern slopes of the Andes, Provincia de Zamora-Chinchipe, República del Ecuador".

Rulyrana mcdiarmidi — Guayasamin, Castroviejo-Fisher, Trueb, Ayarzagüena, Rada, and Vilà, 2009, Zootaxa, 2100: 34.

Common Names

McDiarmid's Glassfrog (Guayasamin, Cisneros-Heredia, McDiarmid, Peña, and Hutter, 2020, Diversity, 12 (222): 215; Coloma and Duellman, 2025, Amph. Ecuador. Vol. 4: xxx).

Rana de Cristal de McDiarmid (Spanish: Guayasamin, Cisneros-Heredia, McDiarmid, Peña, and Hutter, 2020, Diversity, 12 (222): 216; Coloma and Duellman, 2025, Amph. Ecuador. Vol. 4: xxx).

Distribution

Widely distributed in the Utcubamba and central Marañón drainages in Cajamarca and Amazonas, Peru, and southeastern foothills of southern Ecuador (Morona Santiago and Zamora Chinchipe provinces) at elevations between 508 and 1711 m.

Geographic Occurrence

Natural Resident: Ecuador, Peru

Comment

New records, a discussion of the possible conspecific relationship with Rulyrana saxiscandens, and additional life history data were provided by Twomey, Delia, and Castroviejo-Fisher, 2014, Zootaxa, 3851: 48–49. Koch, Venegas, Santa-Cruz Farfan, and Böhme, 2018, Zootaxa, 4385: 25–26, commented on distribution in the Marañon Valley region of Peru. Guayasamin, Cisneros-Heredia, McDiarmid, Peña, and Hutter, 2020, Diversity, 12 (222): 215–217, provided a detailed account, including adult morphology, relationships, natural history, and conservation status and who suggested that Rulyrana mcdiarmidi might be a geographical variant of Rulyrana flavopunctata. Arpi Lojano, Siavichay Pesántez, Aguilar, and Sánchez-Nivicela, 2023, Rev. Latinoam. Herpetol., 6: 21–32, detailed the morphology and developmental stages of the tadpole. J. M. Guayasamin, L. A. Coloma, and A. Terán-Valdez in Coloma and Duellman, 2025, Amph. Ecuador. Vol. 4: 299–301, provided an account, with photographs, which summarized identification, adult morphology, systematics, natural history, distribution (including a dot map for Ecuador), and conservation.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador