Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Vitreorana helenae (Ayarzagüena, 1992)

Class: Amphibia > Order: Anura > Family: Centrolenidae > Subfamily: Centroleninae > Genus: Vitreorana > Species: Vitreorana helenae

Centrolenella helenae Ayarzagüena, 1992, Publ. Asoc. Amigos Doñana, 1: 21. Holotype: MHNLS 9431, by original designation. Type locality: "Quebrada Jaspe, San Ignacio de Yuruaní, Edo Bolívar. Venezuela."

Cochranella helenae — Duellman, 1993, Univ. Kansas Mus. Nat. Hist. Spec. Publ., 21: 45; Cisneros-Heredia and McDiarmid, 2007, Zootaxa, 1572: 1–82.

Hyalinobatrachium helenae — Barrio-Amorós, 1999 "1998", Acta Biol. Venezuelica, 18: 14–15.

Vitreorana helenae — Guayasamin, Castroviejo-Fisher, Trueb, Ayarzagüena, Rada, and Vilà, 2009, Zootaxa, 2100: 39.

English Names

Venezuela Cochran Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 47).

Distribution

Known from the Gran Sabana region of Venezuela (including Quebrada Jaspe, San Ignacio de Yuruaní) as well as Kaieteur National Park, Guyana.

Comment

Very similar to Cochranella oyampiensis (as Centrolenella) according to the original publication. Placed in the Cochranella spinosa group by Savage, 2002, Amph. Rept. Costa Rica: 365. Transferred to Hyalinobatrachium by Barrio-Amorós, 1999 "1998", Acta Biol. Venezuelica, 18: 14–15, but this was rejected by Cisneros-Heredia and McDiarmid, 2007, Zootaxa, 1572: 1–82. Señaris and Ayarzagüena, 2005, Rev. Taxonom. Fam. Centrolenidae de Venezuela: 113–121, provided a review of the morphology, natural history, and range of this species (as Cochranella helenae). See account by Kok and Castroviejo-Fisher, 2008, Zootaxa, 1680: 39–43, who provided new records from Guyana and Venezuela (and noted that the record from Salto Karuay had previously been assigned to Cochranella oyampiensis) and noted this species to be a member of the Cochranella helenae group. Kok and Kalamandeen, 2008, Intr. Taxon. Amph. Kaieteur Natl. Park: 140–141, provided an account. See Cole, Townsend, Reynolds, MacCulloch, and Lathrop, 2013, Proc. Biol. Soc. Washington, 125: 384–385, for brief account and records for Guyana. Señaris, Lampo, Rojas-Runjaic, and Barrio-Amorós, 2014, Guía Ilust. Anf. Parque Nac. Canaima: 84–85, provided a photograph and a brief account for the Parque Nacional de Canaima, Venezuela. See Barrio-Amorós, Rojas-Runjaic, and Señaris, 2019, Amph. Rept. Conserv., 13 (1: e180): 23, for comments on range, taxonomy, and literature. Señaris and Rojas-Runjaic, 2020, in Rull and Carnaval (eds.), Neotrop. Divers. Patterns Process.: 571–632, commented on range and conservation status in the Venezuelan Guayana.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.