Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyloxalus pulchellus (Jiménez de la Espada, 1875)

Class: Amphibia > Order: Anura > Superfamily: Dendrobatoidea > Family: Dendrobatidae > Subfamily: Hyloxalinae > Genus: Hyloxalus > Species: Hyloxalus pulchellus

Phyllodromus pulchellum Jiménez de la Espada, 1875, Vert. Viaje Pacif. Verif. 1862–1865: Pl. 3, fig. 3. Holotype: Frog illustrated in Fig. 3 of the original paper, presumably originally deposited in MNCN but now lost; KU 142976 designated neotype by Coloma, 1995, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 87: 46. Type locality: Not designated; neotype from "2.5 km WNW Cuyuja, 2550 m, Provincia Napo, Ecuador".

Phyllodromus pulchellus — Boulenger, 1882, Cat. Batr. Sal. Coll. Brit. Mus., Ed. 2: 139.

Prostherapis pulchellus — Peracca, 1904, Boll. Mus. Zool. Anat. Comp. Univ. Torino, 19 (465): 17.

Phyllobates taeniatus Andersson, 1945, Ark. Zool., 37A(2): 8. Holotype: NHRM 1904, according to XXX. Type locality: "Rio Cosanga near Archidona, 800 m", Provincia Pastaza, Ecuador. Synonymy by Coloma, 1995, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 87: 46.

Phyllobates riocosangae Andersson, 1945, Ark. Zool., 37A(2): 10. Syntypes: NHRM 1905–06 (2 specimens), according to XXX. Type locality: "Rio Cosanga near Archidona, 600 m", Provincia Pastaza, Ecuador. Synonymy (with Colostethus taeniatus) by Edwards, 1974, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 30: 3.

Phyllobates cosangae — Peters, 1955, Rev. Ecuat. Entomol. Parsitol., 2: 343. Incorrect subsequent spelling.

Colostethus pulchellus — Edwards, 1971, Proc. Biol. Soc. Washington, 84: 148.

Colostethus riocosangae — Edwards, 1971, Proc. Biol. Soc. Washington, 84: 148.

Colostethus taeniatus — Edwards, 1971, Proc. Biol. Soc. Washington, 84: 148.

Colostethus tergogranularis Rivero, 1991, Caribb. J. Sci., 27: 6. Holotype: USNM 282638, by original designation. Type locality: "Faldas, S. of Sumaco, Provincia Napo", Ecuador. Synonymy by Coloma, 1995, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 87: 46.

Hyloxalus pulchellus — Grant, Frost, Caldwell, Gagliardo, Haddad, Kok, Means, Noonan, Schargel, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 299: 169.

Common Names

Espada's Rocket Frog (Colostethus pulchellus: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 49).

Rio Cosanga Rocket Frog (Colostethus taeniatus: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 49).

Faldas Rocket Frog (Colostethus tergogularis: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 49).

Distribution

Elevations of 1590–2970 m in the inter-Andean páramos of northern Ecuador and adjacent Colombia (Nariño and Cauca) and on the eastern slopes of the Andes from southern Colombia south in Ecuador to the headwaters of the Río Pastaza, 1590–2970 m elevation.

Geographic Occurrence

Natural Resident: Colombia, Ecuador

Comment

See account by Coloma, 1995, Misc. Publ. Mus. Nat. Hist. Univ. Kansas, 87: 46. Almendáriz C. and Orcés, 2004, Rev. Politécnica, Quito, 25: 111, provided distributional data for Ecuador. See photograph, map, description of geographic range and habitat, and conservation status (as Colostethus pulchellus) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 225. See note by Guayasamin and Funk, 2009, Zootaxa, 2220: 41–66, on this species at the Yanayacu Biological Station, Napo, Ecuador. Pisso-Florez, Liévano-Bonilla, Mazepa, Mulder, and Ramírez-Chaves, 2023, S. Am. J. Herpetol., 28: 78–88, provided a dot map of this species and Hyloxalus cf. pulchellus and discussed its phylogenetic position.

External links:

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For access to general information see Wikipedia
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For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to Ecuador see FaunaWebEcuador: Anfibios del Ecuador
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.