Minervarya agricola (Jerdon, 1853)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Minervarya > Species: Minervarya agricola

Rana agricola Jerdon, 1853, J. Asiat. Soc. Bengal, 22: 532. Types: In ZSIC, but unavailable for study according to Dubois, 1984, Alytes, 3: 152; reported as lost by Jerdon, 1870, Proc. Asiat. Soc. Bengal, 1870: 67. ZSI/SRS/V/A/362 designated neotype by Ganesh, Dutta, and Chandramouli, 2017, Asian J. Conserv. Biol., 6: 108. Type locality: "inundated paddy-fields and meadows", in South India by implication. Neotype from "Selaiyur Lake in Madras", India. 

Fejervarya granosa Kuramoto, Joshy, Kurabayashi, and Sumida, 2008 "2007", Curr. Herpetol., Kyoto, 26: 94. Holotype: BNHS 4649, by original designation. Type locality: "Talapu, Madikeri", central Western Ghats of Karnataka, India. Synonymy by Chandramouli, Ankaiah, Arul, Dutta, and Ganesh, 2019, Asian J. Conserv. Biol., 8: 85. 

Zakerana granosa — Howlader, 2011, Bangladesh Wildl. Bull., 5: 2.

Fejervarya agricola — Ganesh, Dutta, and Chandramouli, 2017, Asian J. Conserv. Biol., 6: 108. 

Fejervarya granosa —  Dinesh, Vijayakumar, Channakeshavamurthy, Torsekar, Kulkarni, and Shanker, 2015, Zootaxa, 3999: 79. 

Minervarya granosa — Sanchez, Biju, Islam, Hasan, Ohler, Vences, and Kurabayashi, 2018, Salamandra, 54: 115. 

Minervarya agricola — Here (11 July 2018). Provisional placement reflecting the partition of Fejervarya by Sanchez, Biju, Islam, Hasan, Ohler, Vences, and Kurabayashi, 2018, Salamandra, 54: 109–116; Phuge, Patil, Pandit, Kulkarni, Chennakeshavamurthy, Deepak, and Dinesh, 2020, Zootaxa, 4869: 451. 

English Names

Granular Fejervarya (Minervarya granosa [no longer recognized}: Dinesh, Radhakrishnan, Gururaja, and Bhatta, 2009, Rec. Zool. Surv. India, Occas. Pap., 302: 16).

Distribution

Most of India to Nepal, Bangladesh, and southwestern Myanmar; possibly to Bhutan, Thailand, and southwestern China. See comment.

Comment

Considered a junior synonym of Rana vittigera by Theobald, 1868, J. Asiat. Soc. Bengal, 37: 80. Considered a junior synonym of Rana limnocharis by Boulenger, 1890, Fauna Brit. India, Rept. Batr.: 450. Dubois, 1984, Alytes, 3: 152, noted that this species might be based on a juvenile Rana tigerina (now in Hoplobatrachus), but inasmuch as workers were not allowed access to the material the status of the species remained in doubt. Dubois, 1987 "1986", Alytes, 5: 60, considered this to be incertae sedis within Hoplobatrachus or Fejervarya; without discussion. Ganesh, Dutta, and Chandramouli, 2017, Asian J. Conserv. Biol., 6: 107–113, discussed the name and applied it via a neotype to a species in plains of South India, as Fejervarya. These authors also provided a considerable amount of natural history information. Köhler, Mogk, Khaing, and Than, 2019, Vert. Zool., Senckenberg, 69: 190, suggested that the distinctiveness of Minervarya granosa from Minervarya pierrei and Minervarya syhadrensis needs to be confirmed. Chandramouli, Ankaiah, Arul, Dutta, and Ganesh, 2019, Asian J. Conserv. Biol., 8: 84–87, provided mtDNA evidence for the synonymy of Minervarya granosa with Minervarya agricola and discussed the possible status of Minervarya syhadrensis and Minervarya pierrei. See Phuge, Patil, Pandit, Kulkarni, Chennakeshavamurthy, Deepak, and Dinesh, 2020, Zootaxa, 4869: 451–492, for study of call and morphometric variation in the Western Ghats of southern India. Garg and Biju, 2021, Asian Herpetol. Res., 12: 345–370, summarized the systematics (morphology and molecular markers) of this member of the Minervarya agricola group and mapped its distribution. Garg, Sivaperuman, Gokulakrishnan, Gopika, Das, and Biju, 2022, Vert. Zool., Senckenberg, 72: 169–199, discussed and substantiated records for the Andaman Islands, India, and suggested that the species extended into Bhutan, Thailand, and southern China, although specimens or specific localities were not noted. Rahman, Nneji, and Hossain, 2022, J. Asia-Pacific Biodiversity, 15: 147–156, excluded this species from the Bangladesh fauna without discussion. Srinivasulu and Kumar, 2022, J. Threatened Taxa, 14: 21268, reported the species from the state of Telangana, south-central India. 

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