Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nanorana ercepeae (Dubois, 1974)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Nanorana > Species: Nanorana ercepeae

Rana ercepeae Dubois, 1974 "1973", Bull. Soc. Zool. France, 98: 496. Holotype: MNHNP 1974.1091, by original designation. Type locality: "torrent Jiuli Gad, près du village de Takundara, Nord-Ouest de Chainpur, Ouest-Népal (29° 40′ N, 81° 06′ E; altitude 2280 m). "

Rana (Paa) ercepeae — Dubois, 1975, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 3, Zool., 324: 1097; Dubois, 1987 "1986", Alytes, 5: 43.

Paa (Paa) ercepeae — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 320.

Nanorana ercepeae — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 239, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 367.

Chaparana (Paa) ercepeae — Ohler and Dubois, 2006, Zoosystema, 28: 781.

Paa ercepeae — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 26.

Nanorana (Paa) ercepeae — Hofmann, Jablonski, Litvinchuk, Masroor, and Schmidt, 2021, PeerJ, 9 (e11793): 11.

Common Names

Torrent Paa Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 102).

R.C.P.'s Paa Frog (Schleich, Anders, and Kästle, 2002, in Schleich and Kästle (eds.), Amph. Rept. Nepal: 79).

Distribution

Low montane regions, around 3000 m in elevation, western to central Nepal, 2200–2600 m elevation, east to the Kangchenjunga–Singalilla complex, eastern Nepal, on the Sikkim, India, border; photograph identified as this species from Thimphu District, Bhutan; controversially into Sikkim, India (see comment).

Geographic Occurrence

Natural Resident: Bhutan, Nepal

Likely/Controversially Present: India

Comment

Closely related to Nanorana rostandi (as Rana), according to the original publication. See account by Dubois, 1976, Cah. Nepal., Doc., 6: 179–189. Anders, 2002, in Schleich and Kästle (eds.), Amph. Rept. Nepal: 268–270, provided an account for the Nepal population. See statement of geographic range, habitat, and conservation status (as Paa ercepeae) in Stuart, Hoffmann, Chanson, Cox, Berridge, Ramani, and Young, 2008, Threatened Amph. World: 630. Das and Dutta, 2007, Hamadryad, 31: 154–181, noted no larval descriptions in the literature. Not addressed, and therefore not assigned to subgenus, by Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, 107: 13765–13770, and Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, Suppl. Inform., doi:10.1073/pnas.1008415107/-/DCSupplemental: 2. See Shah and Tiwari, 2004, Herpetofauna Nepal: 63, for brief account. Chettri, Bhupathy, and Acharya, 2011, Biodiversity of Sikkim: 233–254, reported the species from Sikkim, India. Subba, Aravind, and Ravikanth, 2016, Check List, 13(1: 2033): 12, considered the presence of this species in Sikkim, India, to be doubtful, although Khambu, 2011, Amph. Kangchenjung–Singalilla Complex, Nepal: ?? [not seen by DRF], reported the species from eastern Nepal on the Sikkim, India, border. Gautam, Chalise, Thapa, and Bhattarai, 2020, IRCF Rept. & Amph., 27: 18–28, briefly discussed abundance and elevational range in the Ghandruk region of central Nepal. Wangyal, Bower, Sherub, Tshewang, Wangdi, Rinchen, Puntsho, Tashi, Koirala, Bhandari, Phuntsho, Koirala, Ghalley, Chaida, Tenzin, Powrel, Tshewang, Raika, Jamtsho, Kinley, Gyeltshen, Tashi, Nidup, Wangdi, Phuentsho, Norbu, Wangdi, Wangchuk, Tobgay, Dorji, and Das, 2020, Herpetol. Rev., 51: 793, identified a photograph of this species from Bhutan. Khatiwada, Wang, Zhao, Xie, and Jiang, 2021, Asian Herpetol. Res., 12: 1–35, did not mention this species as being part of the Nepalese fauna. But, inasmuch as its type locality is in Nepal this exclusion is curious.

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