Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nanorana sichuanensis (Dubois, 1987)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Nanorana > Species: Nanorana sichuanensis

Rana (Paa) sichuanensis Dubois, 1987 "1986", Alytes, 5: 47. Holotype: NHMW 3419.2, by original designation. Type locality: "Ning Yuan Fu (montagne près Xichang, 27° 58′ N, 102° 13′ E, sud du Sichuan, Chine".

Rana muta Su and Li, 1986, Acta Herpetol. Sinica, Chengdu, N.S.,, 5 (2): 152–154. Holotype: KIZ 79006, by original designation. Type locality: "Ninglang County, Yunnan [Province], alt. 2,650 m", China. Preoccupied by Rana muta Laurenti, 1768 (= Rana temporaria Linnaeus, 1758). Synonymy wth Nanorana yunnanensis by Che, Hu, Zhou, Murphy, Papenfuss, Chen, Rao, Li, and Zhang, 2009, Mol. Phylogenet. Evol., 50: 71, and Chen and Hu, 2009, Sichuan J. Zool., 28: 696–699. Synonymy with Rana sichuanensis by Huang, Hu, Wang, Song, Zhou, and Jiang, 2016, Integrative Zool., 11: 144.

Paa (Paa) muta — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 158.

Rana (Paa) liui Dubois, 1987 "1986", Alytes, 5: 150. Replacement name for Rana muta Su and Li, 1986.

Paa (Gynandropaa) liui — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 319.

Rana liui — Zhao and Adler, 1993, Herpetol. China: 144.

Paa (Paa) liui — Fei, 1999, Atlas Amph. China: 216).

Nanorana liui — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 239, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 367.

Gynandropaa liui — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 25. Unexplained recognition.

Paa (Gynandropaa) sichuanensis — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 319.

Nanorana sichuanensis — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 239, by implication.

Gynandropaa (Gynandropaa) sichuanensis — Ohler and Dubois, 2006, Zoosystema, 28: 781. Undiscussed elevation.

Gynandropaa (Gynandropaa) liui — Ohler and Dubois, 2006, Zoosystema, 28: 781.

Gynandropaa sichuanensis — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 25.

Nanorana (Chaparana) sichuanensis — Tang, Liu, and Yu, 2023, Animals, 13(3427): 17.

Common Names

Sichuan Paa Frog (Paa sichuanensis: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 103).

Ninglang Paa Frog (Nanorana liui [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 102).

Vocal Sacless Spiny Frog (Nanorana liui [no longer recognized]: Fei, 1999, Atlas Amph. China: 218).

四川棘蛙 (Chinese: Wang, Ren, Chen, Lyu, Guo, Jiang, Chen, Li, Guo, Wang, and Che, 2020, Biodiversity Sci., 28: App. 1: 11; Yuan, Chen, Wu, Li, and Che, 2022, Biodiversity Sci., 30 (4: 21470): 6).

Distribution

Southern Sichuan into central Yunnan, China.

Geographic Occurrence

Natural Resident: China, People's Republic of

Endemic: China, People's Republic of

Comment

Removed from the synonymy of Nanorana yunnanensis by Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 319, where it had been placed by Inger, Zhao, Shaffer, and Wu, 1990, Fieldiana, Zool., N.S., 58: 12–13; Zhao and Adler, 1993, Herpetol. China: 151. Yang, 1991, Amph. Fauna of Yunnan: 137–138 (as Rana muta), and Fei, 1999, Atlas Amph. China: 218–219 (as Rana liui), provided brief accounts, figures, and maps. See accounts by Yang, 1991, Amph. Fauna of Yunnan: 137 (as Rana muta), and 161–163 (as ; and Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 286; and Fei, 1999, Atlas Amph. China: 219–221. Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 339, noted that this species (Paa liui) may be a junior synonym of Paa sichuanensis. Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 339, noted that this species (Paa liui) may be a junior synonym of Paa sichuanensis. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 186 (as Paa liui) and 187 (Paa sichuanensis), provided brief accounts and illustrations. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1415–1429, provided accounts (as Paa liui and Paa sichuanensis) for China, figures, and map, and included this species in their Paa yunnanensis group. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1415–1421, provided an account (as Paa sichuanensis) for China, figures, and map, and included this species in their Paa yunnanensis group. See Nanorana yunnanensis account for literature under that name that may refer to this species. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 380 (as Paa liui), provided a brief account including photographs of specimens and habitat. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 449, 451, provided accounts (as Gynandropaa liui: 449; and Gynandropaa yunnanensis: 451), photographs, and range maps for China. Not addressed, and therefore not assigned to subgenus, by Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, 107: 13765–13770, and Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, Suppl. Inform., 107(31): 2.Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 376, provided a brief account (as Paa sichuanensis) including photographs. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 450, provided an account, photographs, and a range map for China. Huang, Hu, Wang, Song, Zhou, and Jiang, 2016, Integrative Zool., 11: 134–150, for discussion of systematics.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For access to general information see Wikipedia
For additional sources of general information from other websites search Google
For access to relevant technical literature search Google Scholar
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China