Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Scarthyla Duellman and de Sá, 1988

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Scarthyla

2 species

Scarthyla Duellman and de Sá, 1988, Tropical Zool., 1: 118. Type species: Scarthyla ostinodactyla Duellman and de Sá, 1988 (= Hyla goinorum Bokermann, 1962), by original designation.

Common Names

Madre de Dios Treefrogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 63).

South American Aquatic Treefrogs (Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 14).

Distribution

Upper Amazon Basin, from northern Bolivia to the Iquitos region of Peru, as well as in adjacent Colombia and Brazil; Maracaibo Basin, Venezuela; Caribbean lowlands, Magdalena Valley, and eastern llanos of Colombia.

Comment

Possibly the sister-taxon of Scinax (as Ololygon), according to the original publication and Duellman and Wiens, 1992, Occas. Pap. Mus. Nat. Hist. Univ. Kansas, 151: 1-23. McDiarmid and Altig, 1990 "1989", Alytes, 8: 51-60, suggested on the basis of anatomy that it might be a member of the Scinax rostrata group. In the Dendropsophini of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 90, who placed this taxon as the sister taxon of Pseudis + Lysapsus.

Contained taxa (2 sp.):

Scarthyla goinorum (Bokermann, 1962)

Scarthyla vigilans (Solano, 1971)

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