Heterixalus variabilis (Ahl, 1930)

Class: Amphibia > Order: Anura > Family: Hyperoliidae > Subfamily: Hyperoliinae > Genus: Heterixalus > Species: Heterixalus variabilis

Megalixalus variabilis Ahl, 1930, Mitt. Zool. Mus. Berlin, 16: 526. Syntypes: ZMB (12 specimens), unnumbered in the original publication; including ZMB 7856, 50108-117, a total 11 specimens, according to Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 440. Type locality: "Nossi Faly [Nosy Faly, SW Ambaro Bay, Ambanja District, Diana Region], bei Madagascar".

Heterixalus variabilisLaurent, 1950, Explor. Parc Natl. Albert, Miss. G.F. de Witte (1933–1935), 64: 14, by implication; Bachmann and Blommers-Schlösser, 1975, Zool. Anz., 194: 21; based on specimens of Heterixalus alboguttatus, according to Blommers-Schlösser, 1982, Beaufortia, 32: 9; Glaw and Vences, 1992, Fieldguide Amph. Rept. Madagascar: 278.

English Names

None noted.


Ambanja and Ambilobe, Madagascar.

Geographic Occurrence

Natural Resident: Madagascar

Endemic: Madagascar


See Blommers-Schlösser and Blanc, 1991, Faune de Madagascar, 75: 126. Formerly included in the synonymy of Heterixalus alboguttatus by Blommers-Schlösser, 1982, Beaufortia, 32: 9, and considered a nomen dubium by Guibé, 1978, Bonn. Zool. Monogr., 11: 87, although Glaw and Vences, 1993, Salamandra, 29: 212-230, applied this name to this species. F. Glaw (pers. comm.) suggests that the validity of this name is still uncertain and may be a synonym of Heterixalus tricolor. Glaw and Vences, 1994, Fieldguide Amph. Rept. Madagascar, Ed. 2: 77, discussed the problems with application of this name. Glaw and Vences, 2007, Field Guide Amph. Rept. Madagascar, Ed. 3: 104, provided an account. Tillack, de Ruiter, and Rödel, 2021, Zoosyst. Evol., Berlin, 97: 440, briefly discussed the syntypes and their collector. 

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