Xenophrys zhangi (Ye and Fei, 1992)

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Megophryinae > Genus: Xenophrys > Species: Xenophrys zhangi

Megophrys zhangi Ye and Fei, 1992, Acta Herpetol. Sinica, 1–2: 50–52. Holotype: CIB 750296, by original designation. Type locality: "Zhangmo [=Zhangmu town], Nyanang [= Nyalam County], Xizang, altitude 1000 m", China.

Megophrys (Xenophrys) zhangiDubois and Ohler, 1998, Dumerilia, 4: 14; Mahony, Foley, Biju, and Teeling, 2017, Mol. Biol. Evol., 34: 756. 

Xenophrys zhangiOhler, 2003, Alytes, 21: 23, by implication; Delorme, Dubois, Grosjean, and Ohler, 2006, Alytes, 24: 17; Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 106: 41; Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 39.

Xenophrys (Xenophrys) zhangi — Fei and Ye, 2016, Amph. China, 1: 699. 

English Names

Zhang's Horned Toad (Fei, 1999, Atlas Amph. China: 124; Mahony, Kamei, Teeling, and Biju, 2018, Zootaxa, 4523: 30).

Zhang's Spadefoot Toad (Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 26).


Known from the type locality at Zhangmo, Nyanang, Xizang, China, on the central Nepal border, 1800–2180 m elevation; Kathamandu Valley of Nepal (1462 to 1865 m elevation). 

Geographic Occurrence

Natural Resident: China, People's Republic of, Nepal


Closely related to Megophrys minor according to the original publication. Fei, 1999, Atlas Amph. China: 124, provided a brief account, figure, and map. See comments by Xie and Wang, 2000, Cultum Herpetol. Sinica, 8: 356-370. In the Megophrys parva group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 82. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 469-, provided an account for China, illustration of the holotype, and spot map. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 216, provided a brief account including photographs. Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 26-27, provided an account. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 230–231, provided an account, photographs, and a range map. Fei and Ye, 2016, Amph. China, 1: 699–700, provided an account, photogrphs, and dot map, as Xenophrys caudoprocta. In the Megophrys major group and discussed by Mahony, Kamei, Teeling, and Biju, 2018, Zootaxa, 4523: 30. O'Connell, Aryal, Sherchan, Dhakal, Chaudhary, and Karmacharya, 2019, J. Nat. Hist., London, 53: 1421-1437, discussed the population in the Kathmandu Valley of central-east Nepal. See detailed account for Tibet by Che, Jiang, Yan, and Zhang, 2020, Amph. Rept. Tibet: 120–122, as Xenophrys zhangi. In the Xenophrys major group of Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 9–40. Khatiwada, Wang, Zhao, Xie, and Jiang, 2021, Asian Herpetol. Res., 12: 1–35, discussed the genetics of the species in Nepal. In the Xenophrys major group of Lyu, Qi, Wang, Zhang, Zhao, Zeng, Wan, Yang, Mo, and Wang, 2023, Zool. Res., Kunming, 44: 380–450, who (p. 410–411), provided an account. 

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