Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hylophorbus Macleay, 1878

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Asterophryinae > Genus: Hylophorbus

17 species

Hylophorbus Macleay, 1878, Proc. Linn. Soc. New South Wales, 2: 136. Type species: Hylophorbus rufescens Macleay, 1878, by monotypy.

Metopostira Méhely, 1901, Termés. Füzetek, 24: 189, 238. Type species: Metopostira ocellata Méhely, 1901, by monotypy. Synonymy by Van Kampen, 1919, Bijdr. Dierkd., 21: 53; Zweifel, 1972, Bull. Am. Mus. Nat. Hist., 148: 449.

Common Names

Mawatta Frogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 90).

Distribution

New Guinea.

Comment

Removed from the synonymy of Asterophrys by Zweifel, 1972, Bull. Am. Mus. Nat. Hist., 148: 449, where it had been placed by Parker, 1934, Monogr. Frogs Fam. Microhylidae: 58. Günther, 2001, Russ. J. Herpetol., 8: 81-104, provided cogent accounts and comparisons. Menzies, 2006, Frogs New Guinea & Solomon Is.: 247-252, discussed systematic issues within Hylophorbus and provided brief accounts. Köhler and Günther, 2008, Mol. Phylogenet. Evol., 47: 353-365, suggested on the basis of molecular evidence that Hylophorbus is monophyletic and the sister taxon of a group of "Callulops" including Callulops major. Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 418, recovered Hylophobus as monophyletic and imbedded in a paraphyletic Mantophryne. But, due to the generally chaotic nature of asterophryine generic taxonomy, these authors placed all asterophryine genera for which they had samples into Asterophrys, with the exception of Gastrophrynoides. Ferreira, Kraus, Richards, Oliver, Günther, Trilaksono, Arida, Hamidy, Riyanto, Tjaturadi, Thébaud, Gaucher, and Fouquet, 2023 "2024", Zool. J. Linn. Soc., 202 (paper version of 2024; Zoobank Publication registration of 2023 electronic publication : C205A49A-A66E-466E-8D6D-CBA702798B0A): 1–22, reviewed the phylogenetics of the genus and detected many unnamed species.

Contained taxa (17 sp.):

Hylophorbus atrifasciatus Kraus, 2013

Hylophorbus extimus Zweifel, 1972

Hylophorbus infulatus (Zweifel, 1972)

Hylophorbus lengguru Ferreira, Kraus, Richards, Oliver, Günther, Trilaksono, Arida, Hamidy, Riyanto, Tjaturadi, Thébaud, Gaucher, and Fouquet, 2023

Hylophorbus maculatus Ferreira, Kraus, Richards, Oliver, Günther, Trilaksono, Arida, Hamidy, Riyanto, Tjaturadi, Thébaud, Gaucher, and Fouquet, 2023

Hylophorbus monophonus Ferreira, Kraus, Richards, Oliver, Günther, Trilaksono, Arida, Hamidy, Riyanto, Tjaturadi, Thébaud, Gaucher, and Fouquet, 2023

Hylophorbus myopicus Zweifel, 1972

Hylophorbus nigrinus Günther, 2001

Hylophorbus picoides Günther, 2001

Hylophorbus proekes Kraus and Allison, 2009

Hylophorbus rainerguentheri Richards and Oliver, 2007

Hylophorbus richardsi Günther, 2001

Hylophorbus rufescens Macleay, 1878

Hylophorbus sextus Günther, 2001

Hylophorbus sigridae Günther, Richards, and Dahl, 2014

Hylophorbus tetraphonus Günther, 2001

Hylophorbus wondiwoi Günther, 2001

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