Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Glyphoglossus Gunther, 1869 "1868"

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Microhylinae > Genus: Glyphoglossus

10 species

Glyphoglossus Günther, 1869 "1868", Proc. Zool. Soc. London, 1868: 483. Type species: Glyphoglossus molossus Günther, 1869 "1868", by monotypy.

Calluella Stoliczka, 1872, Proc. Asiat. Soc. Bengal, 1872: 146. Type species: Megalophrys guttulata Blyth, 1856 "1855", by original designation. Synonymy by Peloso, Frost, Richards, Rodrigues, Donnellan, Matsui, Raxworthy, Biju, Lemmon, Lemmon, and Wheeler, 2016, Cladistics, 32: xxx.

Colpoglossus Boulenger, 1904, Ann. Mag. Nat. Hist., Ser. 7, 13: 42. Type species: Colpoglossus brooksi Boulenger, 1904, by monotypy. Synonymy by Parker, 1934, Monogr. Frogs Fam. Microhylidae: 31; Inger, 1966, Fieldiana, Zool., 52: 118.

Dyscophina Van Kampen, 1905, Zool. Jahrb., Jena, Abt. Syst., 22: 708. Type species: Dyscophina volzi Van Kampen, 1905, by monotypy. Synonymy by Parker, 1934, Monogr. Frogs Fam. Microhylidae: 27.

Calliglutus Barbour and Noble, 1916, Proc. New England Zool. Club, 6: 20. Type species: Calliglutus smithi Barbour and Noble, 1916, by monotypy. Synonymy with Colpoglossus by Parker, 1934, Monogr. Frogs Fam. Microhylidae: 30.

Kalluella — Gee and Boring, 1929, Peking Nat. Hist. Bull., 4: 25. Incorrect subsequent spelling.

Common Names

Balloon Frogs (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 89).

Squat Frogs (Calluella [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 88).

Distribution

Southeastern Asia.

Comment

See identification table (as Calluella) by Manthey and Grossmann, 1997, Amph. Rept. Südostasiens: 46, to compare the species of the Sunda Shelf region. Calluella was xcluded from Dyscophinae and placed in Calluellinae by by Fei, Ye, and Jiang, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 278. Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 230, placed Calluella in Microhylinae on the basis of phylogenetic position. See Taylor, 1962, Univ. Kansas Sci. Bull., 43: 575–578 (for Glyphoglossus molossus). Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 225, removed Glyphoglossus (in the sense of not including Calluella) from any subfamily pending resolution of its phylogenetic placement. Transferred to Microhylinae by Van Bocxlaer, Roelants, Biju, Nagaraju, and Bossuyt, 2006, PLoS One, 1: 1–6. Matsui, Hamidy, Belabut, Ahmad, Panha, Sudin, Khonsue, Oh, Yong, Jiang, and Nishikawa, 2011, Mol. Phylogenet. Evol., 61: 167–176, suggested the possibility that this taxon may be imbedded within Calluella on the basis of mtDNA phylogenetic analysis. Pyron and Wiens, 2011, Mol. Phylogenet. Evol., 61: 543–583, in their molecular study, suggested Glyphoglossus to be the sister taxon of Calluella. Das, Min, Hsu, Hertwig, and Haas, 2014, Zootaxa, 3785: 559–560, confirmed the placement of Glyphoglossus within Calluella. Gorin, Solovyeva, Hasan, Okamiya, Karunarathna, Pawangkhanant, de Silva, Juthong, Milto, Nguyen, Suwannapoom, Haas, Bickford, Das, and Poyarkov, 2020, PeerJ, 8 (e9411): 1–47, placed this taxon as the sister of Microhyla.

Contained taxa (10 sp.):

Glyphoglossus brooksii (Boulenger, 1904)

Glyphoglossus capsus (Das, Min, Hsu, Hertwig, and Haas, 2014)

Glyphoglossus flavus (Kiew, 1984)

Glyphoglossus guttulatus (Blyth, 1856)

Glyphoglossus huadianensis Zhang, Liu, Zhang, Hui, Xiao, and Rao, 2021

Glyphoglossus minutus (Das, Yaakob, and Lim, 2004)

Glyphoglossus molossus Günther, 1869

Glyphoglossus smithi (Barbour and Noble, 1916)

Glyphoglossus volzi (Van Kampen, 1905)

Glyphoglossus yunnanensis (Boulenger, 1919)

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