Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Pipa arrabali Izecksohn, 1976

Class: Amphibia > Order: Anura > Family: Pipidae > Subfamily: Pipinae > Genus: Pipa > Species: Pipa arrabali

Pipa arrabali Izecksohn, 1976, Rev. Brasil. Biol., 36: 508. Holotype: EI 5311, by original designation; reported by Peixoto and Gomes, 2007, Rev. Brasil. Zool., 24: 726, as destroyed. Type locality: "Vila Amazônia, Município de Parintins, Estado do Amazonas, Brasil".

English Names

Arrabal's Surinam Toad (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 96).

Arrabal's Pipa (Kok and Kalamandeen, 2008, Intr. Taxon. Amph. Kaieteur Natl. Park: 236).

Distribution

Guyana, western Suriname, eastern Venezuela, and northern (Pará), central (Amazonia and Mato Grosso), and western (Rondônia) Brazil as far east as the state of Tocantins.

Comment

Barrio-Amorós, 1999 "1998", Acta Biol. Venezuelica, 18: 59, commented on the Venezuelan distribution. Zimmerman and Rodrigues, 1990, in Gentry (ed.), Four Neotropical Rainforests: 426–454, provided the first central Brazilan Amazonia record for this species. Sousa and Bernarde, 2007, Herpetol. Rev., 38: 98, provided a record for Rondonia, Brazil. Kok and Kalamandeen, 2008, Intr. Taxon. Amph. Kaieteur Natl. Park: 236–237, provided an account. Señaris, Lampo, Rojas-Runjaic, and Barrio-Amorós, 2014, Guía Ilust. Anf. Parque Nac. Canaima: 236–237, provided a photograph and a brief account for the Parque Nacional de Canaima, Venezuela. Araújo, Haddad, Silva, and Púgener, 2016, An. Acad. Brasil. Cienc., 88: 1875–1887, provided a larval staging table. See Barrio-Amorós, Rojas-Runjaic, and Señaris, 2019, Amph. Rept. Conserv., 13 (1: e180): 109, for comments on range and literature, noting reidentifications and doubting the distinction of this species from Pipa aspera. Silva, Carvalho, Pereira Silva, Fadel, Dantas, Brandão, and Santana, 2020, Biota Neotrop., 20 (1: e20190838): 15, reported the species in Tocantins, Brazil, and discussed the known range in detail. Fouquet, Cornuault, Rodrigues, Werneck, Hrbek, Acosta-Galvis, Massemin, Kok, and Ernst, 2022, Mol. Phylogenet. Evol., 170 (107442): 1–14, suggested on the basis of molecular analysis that nominal Pipa arrabali is composed of at least 3 species, one of which (Guyana) is more closely related to Pipa aspera than to other populations of Pipa arrabali.

External links:

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For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.