Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rana johnsi Smith, 1921

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Rana > Species: Rana johnsi

Rana sauteri var. johnsi Smith, 1921, Proc. Zool. Soc. London, 1921: 434. Syntypes: BMNH (6 specimens—formerly M. Smith 2638-41, 2644, 2657), by original designation. Type locality: "Sui Kat", Langbian Plateau, Vietnam.

Hylarana sauteri johnsi — Bourret, 1939, Annexe Bull. Gen. Instr. Publique, Hanoi, 1939: 59.

Pseudorana sauteri johnsi — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 137; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 233.

Pseudorana johnsi — Jiang, Fei, Ye, Zeng, Zhen, Xie, and Chen, 1997, Cultum Herpetol. Sinica, 6–7: 74; Fei and Ye, 2001, Color Handbook Amph. Sichuan: 190; Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 115; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 37. See comment under Ranidae record.

Rana johnsi — Inger, Orlov, and Darevsky, 1999, Fieldiana, Zool., N.S., 92: 14; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370.

Rana (Rana) johnsi — Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

English Names

John's Groove-toed Frog (Fei, 1999, Atlas Amph. China: 170).

Johns's Frog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 142).

Johns' Frog (Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 28).

Distribution

Known from Jinxiu and Dayaoshan in Guangxi province, and Jianfenglian in Hainan province in China, and from the Lang Bian and Kon Tum Highlands of southern Vietnam through the Annamite mountains of Laos (Bolikhamxai, Khammouan, and Xaignaouli provinces) and Vietnam , and also from Thailand in Phu Luang, Loei Province, 500-700 m elevation, and in Thailand above 1000 m elevation, and in China above 500 m elevation.

Comment

See brief account (as Rana sauteri) by Bourret, 1942, Batr. Indochine: 321-324. Zhao and Adler, 1993, Herpetol. China: 148-149, who suggested that the nominal subspecies (Rana sauteri johnsi in Vietnam and southern China, Rana sauteri sauteri in Taiwan) might be different species. Inger, Orlov, and Darevsky, 1999, Fieldiana, Zool., N.S., 92: 14, subsequently discussed the issue and made the change. Fei, 1999, Atlas Amph. China: 170-171, provided a brief account (as Pseudorana sauteri johnsi), map, and figure. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 190, provided a brief account and illustration (as Pseudorana johnsi). Stuart, 1999, in Duckworth et al. (eds.), Wildlife in Lao PDR: 48, reported the species for the Annamite Mountains of Laos. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 142-143, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 28, provided specific localities for Vietnam and (p. 149) a photograph. Stuart, Sok, and Neang, 2006, Raffles Bull. Zool., 54: 140, provided a record for Cambodia. Stuart, 2005, Herpetol. Rev., 36: 477-478, provided specific localities for Laos. See comment under Pseudorana. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1106-1112, provided an account (as Pseudorana johnsi), figures, and map for China. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 298-299, provided a brief account (as Pseudorana johnsi) including photographs of specimens and habitat. Shi, 2011, Amph. Rept. Fauna Hainan: 104-105, provided an account for Hainan. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 336–337, provided an account (as Pseudorana johnsi), photographs, and a range map for China. Shen, 2014, Fauna Hunan, Amph.: 226–229, provided a detailed account for Hunan, China, as Pseudorana johnsi. Luong, Nguyen, Le, Nguyen, and Nguyen, 2019, Herpetol. Notes, 12: 375–387, provided records from Dien Bien Province, Vietnam, and discussed the range and natural history. Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom, and Poyarkov, 2020, Amph. Rept. Conserv., 14 (2: e248): 218–249, discussed the range and provided a record from Xaignabouli Province, Laos.

External links:

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For additional information see AmphibiaWeb report
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For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.