Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Rana kukunoris Nikolskii, 1918

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Rana > Species: Rana kukunoris

Rana amurensis kukunoris Nikolskii, 1918, Fauna Rossii, Zemnovodnye: 82. Syntypes: ZISP 1500 (3 specimens), by original designation. Type locality: Lake Kukunor [= Qinghai Lake], Qinghai Province, China.

Rana weigoldi Vogt, 1924, Zool. Anz., 60: 339. Syntypes: ZMB (4 specimens), by original designation. Type locality: "Ngo-la [= Ngolo Pass] 4000 m hoher Pass in Derge, Grenzlandschaft im südlichen Osttibet [actually Sichuan] bei Kansego", China. Synonymy with Rana chensinensis by Liu, 1950, Fieldiana, Zool. Mem., 2: 280-281; with Rana kukunoris by Xie, Ye, Fei, and Jiang, 2000, Acta Zootaxon. Sinica, 25: 228-235.

Rana amurensis kokonoris — Liu and Hu, 1961, Tailless Amph. China: 183. Incorrect subsequent spelling.

Rana chensinensis hongyuanensis Hu, Jiang, and Zhao, 1985, Acta Herpetol. Sinica, Chengdu, N.S.,, 4 (3): 228. Types: Not stated, presumably CIB. Type locality: "Mt. Hengduan, 3000-3800 [m elevation]", Sichuan Province, China, 3000-3800 meters. Nomen nudum. Synonymy by Zhao and Adler, 1993, Herpetol. China: 141; with Rana kukunoris by Xie, Ye, Fei, and Jiang, 2000, Acta Zootaxon. Sinica, 25: 228-235.

Rana chensinensis lanzhouensis Wei and Chen, 1990, Acta Zool. Sinica, 36: 78. Types: Not stated, but NWUX 860130, according to Wei, Chen, Xu, and Li, 1991, Acta Zootaxon. Sinica, 16: 381. Type locality: Yuzhong, Sichuan Province, China, 1500 m. Redescribed as new by Wei, Chen, Xu, and Li, 1991, Acta Zootaxon. Sinica, 16: 381. Synonymy with Rana kukunoris by Xie, Ye, Fei, and Jiang, 2000, Acta Zootaxon. Sinica, 25: 228-235.

Rana chensinensis kangdingensis Wei and Chen, 1990, Acta Zool. Sinica, 36: 78. Types: Not stated, but NWUX 860069, according to Wei, Chen, Xu, and Li, 1991, Acta Zootaxon. Sinica, 16: 382. Type locality: Kangding, Sichuan Province, China, 3300 m elevation. Redescribed as new by Wei, Chen, Xu, and Li, 1991, Acta Zootaxon. Sinica, 16: 378. Synonymy with Rana kukunoris by Xie, Ye, Fei, and Jiang, 2000, Acta Zootaxon. Sinica, 25: 228-235.

Rana (Rana) kukunoris — Yuan, Zhou, Chen, Poyarkov, Chen, Jang-Liaw, Chou, Matzke, Iizuka, Min, Kuzmin, Zhang, Cannatella, Hillis, and Che, 2016, Syst. Biol., 65: 835.

English Names

Plateau Brown Frog (Fei and Ye, 2001, Color Handbook Amph. Sichuan: 172; Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 34).

Plateau Wood Frog (Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 34).

Distribution

Plateau region of western China (northwestern Sichuan, eastern Qinghai, Gansu, and very eastern Tibet).

Comment

Removed from the synonymy of Rana chensinensis by Xie, Ye, Fei, and Jiang, 2000, Acta Zootaxon. Sinica, 25: 228-235, where it had been placed by Zhao and Adler, 1993, Herpetol. China: 141. Fei and Ye, 2001, Color Handbook Amph. Sichuan: 172-173, provided a brief account and illustration. In the Rana chensinensis group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 107. Li, Lu, and Li, 2005, Sichuan J. Zool., 24: 268-270, provided a distribution map for China. Che, Pang, Zhao, Matsui, and Zhang, 2007, Zool. Sci., Tokyo, 24: 71-80, discussed that status of this taxon. Li, Lu, Wang, Wang, Fang, and Li, 2005, Herpetol. Sinica, 10: 63-67, compared this species in China with Rana zhenhaiensis, Rana chensinensis, Rana dybowskii, Rana amurensis, and Rana huanrenensis. Zhao, Dai, and Fu, 2009, J. Zool., London, 279: 270-276, reported on landscape genetics in the Hengduan Mountains of Sichuan, China. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1019-1025, provided an account for China and included it in their Rana asiatica group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 274-275, provided a brief account including photographs of specimens and habitat. Li, Zhao, and Dong, 2010, Amph. Rept. Tibet: 34-35, provided an account for Xizang, China. Zhou, Wen, Fu, Xu, Jin, Ding, Min, Che, and Zhang, 2012, Mol. Ecol., 21: 960-973, reported on the phylogenetics and biogeography of this species as part of a larger study of the Rana chensinensis complex. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 310–311, provided an account, photographs, and a range map for China. Feng, Chen, Hu, and Jiang, 2015, Asian Herpetol. Res., 6: 291–297, reported on morphological variation and sexual dimorphism. Wang, Li, Gao, Liu, and Teng, 2020, MtDNA, Part B, 5: 586–587, reported on the complete mitochondrial genome. Che, Jiang, Yan, and Zhang, 2020, Amph. Rept. Tibet: 245–249, provided a detailed account for Tibet, China.

External links:

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For additional sources of information from other sites search Google
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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist; for a quick link to their maps see iNaturalist KML
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.