Zhangixalus feae (Boulenger, 1893)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Zhangixalus > Species: Zhangixalus feae

Rhacophorus feae Boulenger, 1893, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 13: 338. Syntypes: (3 specimens) MSNG 29415, BMNH , and ZMH A04063 (according to Hallermann, 2006, Mitt. Hamburg. Zool. Mus. Inst., 103: 140); MSNG 29415 designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 216. Type locality: "Thao", northern Myanmar.

Rhacophorus (Rhacophorus) feaeAhl, 1931, Das Tierreich, 55: 157; Dubois, 1987 "1986", Alytes, 5: 77.

Rhacophorus nigropalmatus feaeWolf, 1936, Bull. Raffles Mus., 12: 202.

Rhacophorus feaeLiu and Hu, 1961, Tailless Amph. China: 261; Inger, 1966, Fieldiana, Zool., 52: 321; Tian, Jiang, Wu, Hu, Zhao, and Huang, 1986, Handb. Chinese Amph. Rept.: 64.

Polypedates feaeLiem, 1970, Fieldiana, Zool., 57: 98.

Zhangixalus feae — Jiang, Jiang, Ren, Wu, and Li, 2019, Asian Herpetol. Res., 10: 7. 

English Names

Thao Whipping Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 113).

Brown-folded Treefrog (Fei, 1999, Atlas Amph. China: 276).

Fea's Treefrog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 158).

Fea's Large Treefrog (Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom, and Poyarkov, 2020, Amph. Rept. Conserv., 14 (2: e248): 231).


Myanmar through northern Thailand and northern Laos to the northern and central highlands of Vietnam, and limited border areas of southwestern Yunnan, China.


Discussed by Bourret, 1942, Batr. Indochine: 439-441, as Rhacophorus nigropalmatus feae. See Liu and Hu, 1961, Tailless Amph. China: 261-262. Yang, 1991, Amph. Fauna of Yunnan: 203-205; and Fei, 1999, Atlas Amph. China: 276-277, provided accounts. Removed from the synonymy of Rhacophorus nigropalmatus by Inger, 1966, Fieldiana, Zool., 52: 321. In the Rhacophorus (Rhacophorus) dennysii group of Dubois, 1987 "1986", Alytes, 5: 77; see comment under Rhacophorus for dissenting opinion regarding the recognition of this group. Orlov, Lathrop, Murphy, and Ho, 2001, Russ. J. Herpetol., 8: 32-35, provided an account of the Vietnam population and discussed its close similarity to Rhacophorus dennysi. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 96, commented on the Vietnamese distribution. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 158-159, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 37, provided specific localities for Vietnam. Stuart, 2005, Herpetol. Rev., 36: 478, provided a record for Phongsaly Province, Laos. Orlov, Nguyen, and Ho, 2008, Russ. J. Herpetol., 15: 67-84, provided a photograph and a key to differentiate this species from others in Vietnam. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 108-109, provided a brief account for Yunnan, China. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 795-799, provided an account and a spot map for China and assigned this species to their Rhacophorus maximus group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 458, provided a brief account for China including photographs of specimens. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 541, provided an account, photographs, and a range map for China. A record for Thuong Tien Nature Reserve, Hoa Binh Province, in northern Vietnam, reported by Luu, Le, Do, Hoang, Nguyen, Bonkowski, and Ziegler, 2014, Herpetol. Notes, 7: 51–58.  Ziegler, Tran, Nguyen, Perl, Wirk, Kulisch, Lehmann, Rauhaus, Nguyen, Lê, and Vu, 2014, Herpetol. Notes, 7: 185–201, provided a record for Ha Giang Province, Vietnam, and briefly discussed the range. Pham, Nguyen, Ziegler, and Nguyen, 2017, Herpetol. Notes, 10: 379–381, provided records for Son La Province, Vietnam, with notes on morphology and ecology. Youjaroen, Chuaynkern, Chuaynkern, Duengkae, Taksintum, and Phochayavanich, 2018, Russ. J. Herpetol., 25: 165–171, provided a map for Thailand. Kane, Michaels, and Tapley, 2018, Zootaxa, 4504: 138–144, reported on larval development. Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom, and Poyarkov, 2020, Amph. Rept. Conserv., 14 (2: e248): 218–249, briefly discussed the natural history and range and provided records from Oudomzai and Louanghabang provinces, Laos. 

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