Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Zhangixalus feae (Boulenger, 1893)

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Zhangixalus > Species: Zhangixalus feae

Rhacophorus feae Boulenger, 1893, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 13: 338. Syntypes: (3 specimens) MSNG 29415, BMNH 1893.10.9.24 (now renumbered 1947.2.8.35 according to museum records), and ZMH A04063 (according to Hallermann, 2006, Mitt. Hamburg. Zool. Mus. Inst., 103: 140); MSNG 29415 designated lectotype by Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 216. Type locality: "Thao", northern Myanmar.

Rhacophorus (Rhacophorus) feae — Ahl, 1931, Das Tierreich, 55: 157; Dubois, 1987 "1986", Alytes, 5: 77.

Rhacophorus nigropalmatus feae — Wolf, 1936, Bull. Raffles Mus., 12: 202.

Rhacophorus feae — Liu and Hu, 1961, Tailless Amph. China: 261; Inger, 1966, Fieldiana, Zool., 52: 321; Tian, Jiang, Wu, Hu, Zhao, and Huang, 1986, Handb. Chinese Amph. Rept.: 64.

Polypedates feae — Liem, 1970, Fieldiana, Zool., 57: 98.

Zhangixalus feae — Jiang, Jiang, Ren, Wu, and Li, 2019, Asian Herpetol. Res., 10: 7.

Rhacophorus (Zhangixalus) feae — Mahony, Kamei, Brown, and Chan, 2024, Vert. Zool., Senckenberg, 74: 253, by implication.

Common Names

Thao Whipping Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 113).

Brown-folded Treefrog (Fei, 1999, Atlas Amph. China: 276).

Fea's Treefrog (Chan-ard, 2003, Photograph. Guide Amph. Thailand: 158).

Fea's Tree Frog (Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 460).

Fea's Large Treefrog (Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom, and Poyarkov, 2020, Amph. Rept. Conserv., 14 (2: e248): 231).

Fea's Gliding Treefrog (Zug, 2022, Smithson. Contrib. Zool., 653: 47).

Distribution

Myanmar (Kayah) through northern Thailand (Mae Hong Song, Chiang Mai, and Loei provinces) and Laos (Xiengkhouang, Louangphabang, and Xekong provinces) to the northern and central highlands of Vietnam, and limited border areas of southwestern Yunnan, China; possibly to Bangladesh, Cambodia, and easternmost India.

Geographic Occurrence

Natural Resident: China, People's Republic of, Laos, Myanmar, Thailand, Vietnam

Likely/Controversially Present: Bangladesh, Cambodia, India

Comment

Discussed by Bourret, 1942, Batr. Indochine: 439–441, as Rhacophorus nigropalmatus feae. See Liu and Hu, 1961, Tailless Amph. China: 261–262. Yang, 1991, Amph. Fauna of Yunnan: 203–205; and Fei, 1999, Atlas Amph. China: 276–277, provided accounts. Removed from the synonymy of Rhacophorus nigropalmatus by Inger, 1966, Fieldiana, Zool., 52: 321. In the Rhacophorus (Rhacophorus) dennysii group of Dubois, 1987 "1986", Alytes, 5: 77; see comment under Rhacophorus for dissenting opinion regarding the recognition of this group. Li and Hu, 1996, Zool. Res., Kunming, 17: 483–488, reported on karyology. Orlov, Lathrop, Murphy, and Ho, 2001, Russ. J. Herpetol., 8: 32–35, provided an account of the Vietnam population and discussed its close similarity to Rhacophorus dennysi. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 96, commented on the Vietnamese distribution. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 158–159, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 37, provided specific localities for Vietnam. Stuart, 2005, Herpetol. Rev., 36: 478, provided a record for Phongsaly Province, Laos. Orlov, Nguyen, and Ho, 2008, Russ. J. Herpetol., 15: 67–84, provided a photograph and a key to differentiate this species from others in Vietnam. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 108–109, provided a brief account for Yunnan, China. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 795–799, provided an account and a spot map for China and assigned this species to their Rhacophorus maximus group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 458, provided a brief account for China including photographs of specimens. Chan-ard, Cota, and Makchai, 2011, Amph. E. Region Thailand: 140, detailed the range in Thailand. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 541, provided an account, photographs, and a range map for China. A record for Thuong Tien Nature Reserve, Hoa Binh Province, in northern Vietnam, reported by Luu, Le, Do, Hoang, Nguyen, Bonkowski, and Ziegler, 2014, Herpetol. Notes, 7: 51–58. Ziegler, Tran, Nguyen, Perl, Wirk, Kulisch, Lehmann, Rauhaus, Nguyen, Le, and Vu, 2014, Herpetol. Notes, 7: 185–201, provided a record for Ha Giang Province, Vietnam, and briefly discussed the range. Pham, Nguyen, Ziegler, and Nguyen, 2017, Herpetol. Notes, 10: 379–381, provided records for Son La Province, Vietnam, with notes on morphology and ecology. Youjaroen, Chuaynkern, Chuaynkern, Duengkae, Taksintum, and Phochayavanich, 2018, Russ. J. Herpetol., 25: 165–171, provided a map for Thailand. Kane, Michaels, and Tapley, 2018, Zootaxa, 4504: 138–144, reported on larval development. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 460–461, provided a brief account (description, photographs, habitat, and range) for Thailand (in Thai). Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 178–179, provided photographs, a brief account for Thailand, and a range map. Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom, and Poyarkov, 2020, Amph. Rept. Conserv., 14 (2: e248): 218–249, briefly discussed the natural history and range and provided records from Oudomzai and Louanghabang provinces, Laos. Zug, 2022, Smithson. Contrib. Zool., 653: 47, briefly discussed identification, habitat, and range in Myanmar. Brakels, Maury, and Nguyen, 2021, Herpetol. Rev., 52: 793, provided a record from Laos, Xiengkhouang Province, Mork District and discussed the range. Pham, Nguyen, Tran, Le, Nguyen, Nguyen, Pham, Dang, and Doan, 2023, VNU J. Sci. Earth Environ. Sci., 39: 53–61, provided a record for Son La Province, Vietnam.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.