Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Nanorana feae (Boulenger, 1887)

Class: Amphibia > Order: Anura > Family: Dicroglossidae > Subfamily: Dicroglossinae > Genus: Nanorana > Species: Nanorana feae

Rana feae Boulenger, 1887, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 5: 418. Holotype: MSNG 29301, according to Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 215. Type locality: "Kakhien Hills", Myanmar; Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 215, noted holotype data as Bhamò, Kakhyen Hills, Myanmar. Biju, Garg, Gokulakrishnan, Sivaperuman, Thammachoti, Ren, Gopika, Bisht, Hamidy, and Shouche, 2020, Zootaxa, 4878: 31, discussed the location of Bhamò.

Rana (Rana) feae — Boulenger, 1920, Rec. Indian Mus., 20: 8.

Rana (Paa) feae — Dubois, 1975, Bull. Mus. Natl. Hist. Nat. Paris, Ser. 3, Zool., 324: 1097; Dubois, 1976, Cah. Nepal., Doc., 6: 24; Dubois, 1987 "1986", Alytes, 5: 43.

Paa (Paa) feae — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 157; Fei, 1999, Atlas Amph. China: 218-219.

Paa (Gynandropaa) feae — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 319.

Nanorana feae — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 239, by implication; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 367.

Gynandropaa (Gynandropaa) feae — Ohler and Dubois, 2006, Zoosystema, 28: 781.

Gynandropaa feae — Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 25.

Common Names

Kakhien Paa Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 102).

Ocellated Spiny Frog (Fei, 1999, Atlas Amph. China: 218).

Fei's Mountain Frog (Zug, 2022, Smithson. Contrib. Zool., 653: 24).

Distribution

Kakhyen Hills in Kachin, Myanmar.

Geographic Occurrence

Natural Resident: Myanmar

Endemic: Myanmar

Comment

See also Dubois, 1987 "1986", Alytes, 5: 43, who discussed this species. Accounts provided by Boulenger, 1920, Rec. Indian Mus., 20: 68; Liu, 1950, Fieldiana, Zool. Mem., 2: 277-278. Yang, 1991, Amph. Fauna of Yunnan: 122-123; and Fei, 1999, Atlas Amph. China: 218-219, provided a brief account, map, and figure.Not addressed, and therefore not assigned to subgenus, by Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, 107: 13765-13770, and Che, Zhou, Hu, Papenfuss, Wake, and Zhang, 2010, Proc. Natl. Acad. Sci. USA, Suppl. Inform., doi:10.1073/pnas.1008415107/-/DCSupplemental: 2. Huang, Hu, Wang, Song, Zhou, and Jiang, 2016, Integrative Zool., 11: 145, suggested that the earlier record from Husa, Yunnan, China, was based on a misidentified Nanorana yunnanensis. Zug, 2022, Smithson. Contrib. Zool., 653: 23, briefly discussed identification, habitat, and range in Myanmar. Tang, Liu, and Yu, 2023, Animals, 13(3427): 17, suggested that this species is unassignable to subgenus.

External links:

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To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.