Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Gastrophryne Fitzinger, 1843

Class: Amphibia > Order: Anura > Family: Microhylidae > Subfamily: Gastrophryninae > Genus: Gastrophryne

4 species

Gastrophryne Fitzinger, 1843, Syst. Rept.: 33. Type species: Engystoma rugosum Duméril and Bibron, 1841 (= Engystoma carolinense Holbrook, 1836), by original designation.

English Names

Narrowmouth Toads (Liner, 1994, Herpetol. Circ., 23: 22; Lee, 2000, Field Guide Amph. Rept. Maya World: 121; Collins and Taggart, 2009, Standard Common Curr. Sci. Names N. Am. Amph. Turtles Rept. Crocodil., ed. 6: 8).

American Narrowmouth Toads (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 89).

North American Narrow-mouthed Toads (Crother, Boundy, Campbell, de Queiroz, Frost, Highton, Iverson, Meylan, Reeder, Seidel, Sites, Taggart, Tilley, and Wake, 2001 "2000", Herpetol. Circ., 29: 9; Frost, McDiarmid, and Mendelson, 2008, in Crother (ed.), Herpetol. Circ., 37: 5; Liner and Casas-Andreu, 2008, Herpetol. Circ., 38: 14; Frost, McDiarmid, Mendelson, and Green, 2012, in Crother (ed.), Herpetol. Circ., 39: 15; Frost, Lemmon, McDiarmid, and Mendelson, 2017, in Crother (ed.), Herpetol. Circ., 43: 10; Hedges, Powell, Henderson, Hanson, and Murphy, 2019, Caribb. Herpetol., 67: 15).

Distribution

Southern USA to Honduras.

Comment

Reviewed by Nelson, 1972, J. Herpetol., 6: 111-137; and Nelson, 1973, Cat. Am. Amph. Rept., 134: 1-2. Probably derived from Hypopachus according to Carvalho, 1954, Occas. Pap. Mus. Zool. Univ. Michigan, 555: 12. A close affinity between these two genera and the lack of close relationship between Gastrophryne and Microhyla (contra Parker, 1934, Monogr. Frogs Fam. Microhylidae) was supported by the subsequent studies summarized by Bogart and Nelson, 1976, Herpetologica, 32: 205. Greenbaum, Smith, and de Sá, 2011, Mol. Phylogenet. Evol., 61: 265-277, suggested that Gastrophryne is paraphyletic with respect to Hypopachus, with Gastrophryne usta being more closely related to Hypopachus than to other Gastrophryne.

Contained taxa (4 sp.):

Gastrophryne carolinensis (Holbrook, 1835)

Gastrophryne elegans (Boulenger, 1882)

Gastrophryne mazatlanensis (Taylor, 1943)

Gastrophryne olivacea (Hallowell, 1856)

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