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Hylarana lateralis (Boulenger, 1887)
Rana lateralis Boulenger, 1887, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 2, 5: 483. Holotype: MSNG 29324, according to Capocaccia, 1957, Ann. Mus. Civ. Stor. Nat. Genova, Ser. 3, 69: 215. Type locality: "Kaw-ka-riet" (=Kokarit), east of Moulmein, Tenasserim, Myanmar.
Rana (Rana) lateralis — Boulenger, 1920, Rec. Indian Mus., 20: 9; Dubois, 1987 "1986", Alytes, 5: 42.
Rana nigrolineata Liu and Hu, 1960 "1959", Acta Zool. Sinica, 11: 516, 530. Holotype: CIB 571085. Type locality: "Meng-yang, Yunnan, 680 meters altitude", China. Synonymy by Ohler, 2007, Alytes, 25: 64.
Rana (Rana) nigrolineata — Dubois, 1987 "1986", Alytes, 5: 42.
Pelophylax nigrolineatus — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 136; Fei, 1999, Atlas Amph. China: 164; Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 369; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1–13; by implication.
Rana (Pelophylax) lateralis — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 332.
Rana (Pelophylax) nigrolineata — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 332.
Hylarana lateralis — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 237, by implication.
Hylarana nigrolineata — Chen, Murphy, Lathrop, Ngo, Orlov, Ho, and Somorjai, 2005, Herpetol. J., 15: 237, by implication.
Pelophylax lateralis — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 369; Che, Pang, Zhao, Wu, Zhao, and Zhang, 2007, Mol. Phylogenet. Evol., 43: 1–13; by implication.
Humerana lateralis — Grosjean, Ohler, Chuaynkern, Cruaud, and Hassanin, 2015, C. R. Biol., Paris, 338: 353.
Hylarana (Humerana) lateralis — Wu, Xu, Suwannapoom, Nguyen, Murphy, Papenfuss, Lathrop, Kilunda, Gao, Yuan, Chen, Zhang, Zhao, Wang, Rahman, Nneji, Zhao, Wang, Jin, Zhang, and Che, 2024 "2025", Mol. Phylogenet. Evol., 108223: 1–11, by implication.
Common Names
Kokarit Frog (Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 39).
Kokarit Frog (Pelophylax lateralis: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 108).
Yellow Frog (Pelophylax lateralis: Chan-ard, 2003, Photograph. Guide Amph. Thailand: 142).
Meng-yang Frog (Pelophylax nigrolineatus [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 108).
Black-lined Pond Frog (Pelophylax nigrolineatus [no longer recognized]: Fei, 1999, Atlas Amph. China: 164).
Kokarit Copper-backed Frog (Zug, 2022, Smithson. Contrib. Zool., 653: 37).
Distribution
Southern Yunnan, China, on the Vietnam and Laos borders; non-peninsular Thailand; southern Myanmar; Cambodia, southern Laos and Vietnam (Ninh Binh, Dong Nai, Lam Dong, Dac Lac, and Gia Lai provinces).
Geographic Occurrence
Natural Resident: Cambodia, China, People's Republic of, Laos, Myanmar, Thailand, Vietnam
Comment
See Taylor, 1962, Univ. Kansas Sci. Bull., 43: 443–446, and Bourret, 1942, Batr. Indochine: 302–304, for discussion. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 89, commented on the range. Nutphund, 2001, Amph. Thailand: 120, provided a brief characterization and photograph, which Ohler, 2003, Alytes, 21: 102, allocated to Limnonectes sp. Stuart, 1999, in Duckworth et al. (eds.), Wildlife in Lao PDR: 48, commented briefly on the Laos range. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 142–143, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 28, provided specific localities for Vietnam. Stuart, 2005, Herpetol. Rev., 36: 478, provided specific localities for Laos. Liu and Hu, 1961, Tailless Amph. China: 175–177, Yang, 1991, Amph. Fauna of Yunnan: 139–140, and Fei, 1999, Atlas Amph. China: 164–166, provided brief accounts as Pelophylax nigrolineatus. In the Pelophylax pleuraden group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 111. See discussion by Ohler, 2007, Alytes, 25: 55–74, who placed Rana nigrolineata into synonymy on the basis of morphological characteristics. Neang and Holden, 2008, Field Guide Amph. Cambodia: 78–79, provided photographs, brief account of identification, ecology, and range in Cambodia. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 69–70, provided a brief account (as Rana nigrolineata). Stuart, 2008, Mol. Phylogenet. Evol., 46: 49–60, suggested on the basis of molecular evidence that this species is imbedded within Hylarana, and far from Pelophylax, but did not make the taxonomic change. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 3: 1075–1080, provided an account (as Pelophylax nigrolineatus), figures, and map for China and included it in their Pelophylax nigromaculatus group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 283 (as Pelophylax nigrolineatus), provided a brief account including photographs. Chan-ard, Cota, and Makchai, 2011, Amph. E. Region Thailand: 112–113, provided a photograph and brief account for the Eastern Region of Thailand, and (p. 146) detailed the range in Thailand. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 327, provided an account (as Pelophylax nigrolineatus), photographs, and a range map. Oliver, Prendini, Kraus, and Raxworthy, 2015, Mol. Phylogenet. Evol., 90: 188, noted that this species is likely in either in Hydrophylax or Indosylvirana, but pending genetic sampling declined to make any taxonomic changes. Subsequently, Grosjean, Ohler, Chuaynkern, Cruaud, and Hassanin, 2015, C. R. Biol., Paris, 338: 351–361, reported on DNA barcoding of larvae and provided the data to make the assignment to Humerana. See account, photograph, and map for Vietnam in Vassilieva, Galoyan, Poyarkov, and Geissler, 2016, Photograph. Field Guide Amph. Rept. Lowland S. Vietnam: 100–102. Niyomwan, Srisom, and Pawangkhanant, 2019, Field Guide Amph. Thailand: 344–345, provided a brief account (description, photographs, habitat, and range) for Thailand (in Thai). Vassilieva, 2019, Zootaxa, 46: 138–140, reported on larval morphology from specimens from Cat Tien National Park, Tan Phu District, Dong Nai Province, Vietnam. Makchai, Chuaynkern, Safoowong, Chuachat, and Cota, 2020, Amph. N. Thailand: 146, provided a brief account, photographs, and a range map for Thailand. See comments by Geissler, Hartmann, Ihlow, Neang, Seng, Wagner, and Böhme, 2019, Cambodian J. Nat. Hist., 2019: 40–63, on specimens collected in Phnom Kulen National Park, northern Cambodia. Zug, 2022, Smithson. Contrib. Zool., 653: 37, briefly discussed identification, habitat, and range in Myanmar. Stuart, Seateun, Sivongxay, and Phimmachak, 2025 "2024", in Wildlife Lao PDR: 39, summarized literature, taxonomy, habitat, and detailed range for Laos.
External links:
Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.
- For access to general information see Wikipedia
- For additional sources of general information from other websites search Google
- For access to relevant technical literature search Google Scholar
- For images search CalPhoto Images and Google Images
- To search the NIH genetic sequence database, see GenBank
- For additional information see AmphibiaWeb report
- For information on conservation status and distribution see the IUCN Redlist
- For information on distribution, habitat, and conservation see the Map of Life
- For related information on conservation and images as well as observations see iNaturalist
- For additional information specific to China see Amphibia China