Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Sanguirana sanguinea (Boettger, 1893)

Class: Amphibia > Order: Anura > Family: Ranidae > Genus: Sanguirana > Species: Sanguirana sanguinea

Rana sanguinea Boettger, 1893, Zool. Anz., 16: 364. Holotype: SMF 6221 (formerly 1062.a) according to Mertens, 1967, Senckenb. Biol., 48(A): 45; MCZ 9935–38 were apparently considered syntypes in error by Barbour and Loveridge, 1929, Bull. Mus. Comp. Zool., 69: 331. Type locality: "Insel Culion, Calamianes-Gruppe, Philippinen".

Rana varians Boulenger, 1894, Ann. Mag. Nat. Hist., Ser. 6, 14: 86. Syntypes: BMNH (3 specimens), by original designation, by museum records these including BMNH 1947.2.29.52–53. Type locality: "Palawan", Philippines. Taylor, 1920, Philipp. J. Sci., 16: 220, and Inger, 1954, Fieldiana, Zool., 33: 328, and Zhao and Adler, 1993, Herpetol. China: 132, but considered a distinct species by Liu and Hu, 1961, Tailless Amph. China: 201–203, and Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 341, and who also recognized Rana nigrotympanica for the Chinese populations formerly recognized under the name Rana varians.

Rana (Hylorana) sanguinea — Boulenger, 1920, Rec. Indian Mus., 20: 127–130.

Rana (Hylorana) varians — Boulenger, 1920, Rec. Indian Mus., 20: 127–130.

Hylorana varians — Deckert, 1938, Sitzungsber. Ges. Naturforsch. Freunde Berlin, 1938: 144.

Rana (Hylarana) sanguinea — Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Rana (Hylarana) varians — Dubois, 1987 "1986", Alytes, 5: 42, by implication.

Hylarana (Hylarana) varians — Fei, Ye, and Huang, 1990, Key to Chinese Amph.: 139. Based on specimens of Hylarana nigrotympanica according to Stuart, Heatwole, and Tan, 2006, Hamadryad, 30: 109.

Rana (Sanguirana) sanguinea — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 329.

Rana (Sanguirana) varians — Dubois, 1992, Bull. Mens. Soc. Linn. Lyon, 61: 329.

Hylarana sanguinea — Song, Jang, Zou, and Shi, 2002, Herpetol. Sinica, 9: 71.

Sanguirana sanguinea — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 37. See comment under Ranidae record.

Sanguirana varians — Frost, Grant, Faivovich, Bain, Haas, Haddad, de Sá, Channing, Wilkinson, Donnellan, Raxworthy, Campbell, Blotto, Moler, Drewes, Nussbaum, Lynch, Green, and Wheeler, 2006, Bull. Am. Mus. Nat. Hist., 297: 370; Fei, Ye, and Jiang, 2010, Herpetol. Sinica, 12: 37. See comment under Ranidae record.

Common Names

Calamianes Frog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 109).

Palawan Frog (Sanguirana varians [no longer recognized]: Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 110).

Distribution

Palawan I., Philippines; Moluccas and Sulawesi, Indonesia; Ceram Island, South Moluccas. See comment.

Geographic Occurrence

Natural Resident: Indonesia, Philippines, South Moluccas

Comment

See accounts by Boulenger, 1920, Rec. Indian Mus., 20: 158; Inger, 1954, Fieldiana, Zool., 33: 328–331. See comment under Hylarana nigrotympanica. Song, Jang, Zou, and Shi, 2002, Herpetol. Sinica, 9: 69–79, reported this species for Hainan I., China, but this is likely a misidentification, probably based on Hylarana nigrotympanica. Identifications outside of Palawan, Philippines, are suspect.

External links:

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For access to general information see Wikipedia
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For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For information on distribution, habitat, and conservation see the Map of Life
For related information on conservation and images as well as observations see iNaturalist
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.