Rhacophorus kio Ohler and Delorme, 2006

Class: Amphibia > Order: Anura > Family: Rhacophoridae > Subfamily: Rhacophorinae > Genus: Rhacophorus > Species: Rhacophorus kio

Rhacophorus kio Ohler and Delorme, 2006, C. R. Biol., Paris, 329: 90. Holotype: MNHNP 2004.0411, by original designation. Type locality: "Long Nai, Nam Lan Forest Conservation Area, Buon Tai County, Phongsaly Province, Laos".

English Names

Black-webbed Treefrog (Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 461).

Asian Black-webbed Treefrog (Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 65). 

Kio Flying Frog (Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom, and Poyarkov, 2020, Amph. Rept. Conserv., 14 (2: e248): 228).


Southern Yunnan and southern Guangxi, China, as well as northern Thailand, eastern India, Laos, Vietnam, and Cambodia; possibly in adjacent Myanmar.


Confused with Rhacophorus reinwardtii prior to its description so all literature prior to 2006 applying to this species refers to it as Rhacophorus reinwardtii. By implication in the Rhacophorus (Rhacophorus) reinwardtii group of Dubois, 1987 "1986", Alytes, 5: 77; see comment under Rhacophorus for dissenting opinion regarding the recognition of this group. For accounts (as Rhacophorus reinwardtii) see Liu and Hu, 1961, Tailless Amph. China: 255–257; Yang, 1991, Amph. Fauna of Yunnan: 218–220; Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 335; and Fei, 1999, Atlas Amph. China: 268–269; Zhang and Wen, 2000, Amph. Guangxi: 133–134; see also Orlov, Lathrop, Murphy, and Ho, 2001, Russ. J. Herpetol., 8: 38–39, who noted that previous records of Rhacophorus nigropalmatus from Vietnam likely applied to this species. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 97, noted that the elevation range in Vietnam is 200–1800 m. Sarkar, Biswas, and Ray, 1992, State Fauna Ser., 3: 93, provided a brief account as Rhacophorus reinwardti, for West Bengal, India. Chan-ard, 2003, Photograph. Guide Amph. Thailand: 162–163, provided a very brief account, map for Thailand, and photograph. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 40, provided specific localities for Vietnam. Bain, Nguyen, and Doan, 2007, Herpetol. Rev., 38: 111, provided records for Thua Thien-Hue Province, Vietnam, treated the previous records of Rhacophorus reinwardtii from Laos as this species, and briefly discussed the range. Orlov, Nguyen, and Ho, 2008, Russ. J. Herpetol., 15: 67–84, provided a photograph and a key to differentiate this species from others in Vietnam. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 774–777, provided an account and a spot map for China and assigned this species to their Rhacophorus verrucopus group. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 461, provided a brief account for China including photographs of specimens. Li, Li, Murphy, Rao, and Zhang, 2012, Zool. Scripta, 41: xxx, provided evidence that all specimens of nominal Rhacophorus reinwardtii from China are referrable to Rhacophorus kio. Yang, 2008, in Yang and Rao (ed.), Amph. Rept. Yunnan: 116–117, provided a brief account (as Rhacophorus reinwardtii) for Yunnan, China. Rowley, Tran, Hoang, and Le, 2012, J. Herpetol., 46: 480–487, provided some comparative molecular data with Rhacophorus helenaeFei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 545, provided an account, photographs, and a range map for China. A record for Thuong Tien Nature Reserve, Hoa Binh Province, in northern Vietnam, reported by Luu, Le, Do, Hoang, Nguyen, Bonkowski, and Ziegler, 2014, Herpetol. Notes, 7: 51–58. Pham, Pham, Hoang, Ziegler, and Nguyen, Herpetol. Notes, 10: 183–191, provided records from Ha Giang Province, Vietnam, as well as morphological and ecological observation. Pham, An, Herbst, Bonkowski, Ziegler, and Nguyen, 2017, Bonn Zool. Bull., 66: 37–53, provided records for Cao Bang Province, Vietnam, along with observatons on morphology and natural history. Reported from Lai Chau Province, Vietnam, by Pham, Hoang, Pham, and Nguyen, VNU J. Sci. Nat. Sci. Tech., 35: 73–79. Zug and Mulcahy, 2020 "2019", Amph. Rept. S. Tanintharyi: 65, provided a brief account for South Tanintharyi, peninsular Myanmar, although the authors give the species' range as excluding peninsular Myanmar. Nguyen, Brakels, Maury, Sudavanh, Pawangkhanant, Idiiatullina, Lorphengsy, Inkhavilay, Suwannapoom, and Poyarkov, 2020, Amph. Rept. Conserv., 14 (2: e248): 218–249, briefly discussed the natural history and range and provided records for Vientiane and Xiasomboun Provinces, Laos.  

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