Hyla simplex Boettger, 1901

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Hyla > Species: Hyla simplex

Hyla chinensis var. simplex Boettger, 1901, Ber. Senckenb. Naturforsch. Ges., 1901 (Wiss. Abhandl.): 53. Holotype: SMF 2626, according to Mertens, 1967, Senckenb. Biol., 48(A): 41. Type locality: "Phuc-Son in Annam", Vietnam.

Hyla simplexBoulenger, 1903, Ann. Mag. Nat. Hist., Ser. 7, 12: 186; Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 477-480; Tian, Jiang, Wu, Hu, Zhao, and Huang, 1986, Handb. Chinese Amph. Rept.: 58.

Hyla chinensis simplexNieden, 1923, Das Tierreich, 46: 201; Bourret, 1937, Annexe Bull. Gen. Instr. Publique, Hanoi, 1937: 21.

Hyla simplex simplexFei and Ye, 2000, Cultum Herpetol. Sinica, 8: 71, by implication.

Hyla simplex hainanensis Fei and Ye, 2000, Cultum Herpetol. Sinica, 8: 71, 73. Holotype: CIB 64III4273, by original designation. Type locality: "Wenchang County (110° 70′ E, 19° 62′ N), Hainan Province, altitude 20 m", China.

Hyla (Hyla) simplex — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331, by implication. 

English Names

Annam Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 58).

South China Tree Toad (Fei, 1999, Atlas Amph. China: 142).

South China Tree Toad (Hyla simplex simplex: Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 256).

Hainan Tree Toad (Hyla simplex hainanensis: Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 256


Vietnam (Annam and Tonkin), northeastern Laos, and southern China (eastern Jiangxi, Zhejiang, Guangxi, western Guangdong, and Hainan), 50 to 1500 m elevation. See comment. 

Geographic Occurrence

Natural Resident: China, People's Republic of, Laos, Vietnam


In the Hyla arborea group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102 (see comment under Hyla), although it is likely to be a member of the Hyla eximia group (DRF). See Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 477–480, Liu and Hu, 1961, Tailless Amph. China: 128–130, and (as Hyla chinensis simplex) Bourret, 1942, Batr. Indochine: 224–226. See account by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 205. Fei, 1999, Atlas Amph. China: 142–144, provided a brief account, map, and figure. Huang, 1990, Fauna Zhejiang, Amph. Rept.: 53–54, provided an account for Zhejiang populations. Zhang and Wen, 2000, Amph. Guangxi: 66, provided an account for Guangxi. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 83, provided the Laos component of the range statement. Zhang, 2002, Sichuan J. Zool., 21: 198–199, provided a key to differentiate this species from others in China. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 18, provided specific localities for Vietnam. In the Hyla immaculata group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 98, who provided a key to the subspecies. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 605–613, provided accounts by species and subspecies, illustrations of holotypes, and spot maps. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 256, provided brief accounts as Hyla simplex simplex and Hyla simplex hainanensis. Shi, 2011, Amph. Rept. Fauna Hainan: 45–46, provided an account for Hainan. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 288–289, provided accounts (as Hyla simplex simplex and Hyla simplex hainanensis), illustrations, and a range map. Fei and Ye, 2016, Amph. China, 1: 854–859, provided accounts by subspecies, photographs, and spot maps for China. Li, 2011, Amph. Rept. Guangdong: 37, provided a brief account for Guangdong, China, and photograph. Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 28, suggested without providing evidence that records from China may actually correspond to Hyla zhaopingensis and that the Laos record requires confirmation.       

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