Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Search tool	Example of search string in box	What that means to the database
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	Hyl*	Will find all uses of "Hyl . . ." anywhere in a record: e.g., Hylarana, Hyla, Hylidae, Hylinae, Hylaedactyla.
* = "wildcard". Allows a partial letter string to be entered and implies either no or various extension on this letter string	*Hyla	Will find all uses of ". . . hyla" anywhere in a record: e.g., Hyla, Hylidae, Plectrohyla, Ptychadena hylaea, Adenomera hylaedactyla
Single word	Hyla	Will find all records that contain stand-alone uses of Hyla: e.g., Hyla, Hyla arenicolor
Strings of words without quotation marks	Lithobates pipiens	Interprets this as "Lithobates OR pipiens" so will find the union of all records that contain either "Lithobates" or "pipiens": e.g., Lithobates omiltemanus, Hylorana pipiens
Strings of word with quotation marks	"Lithobates pipiens"	Interprets this as "Lithobates AND pipiens" so will return all records that have the character string "Lithobates pipiens" anywhere within a record: e.g., all members of the Lithobates pipiens complex.
AND statement. Means that the results of the search must include both words held together by an AND	Lithobates AND pipiens	Same as "Lithobates pipiens"
OR statement. Means that the results of the search will all records that have either of the terms held together by an OR	Lithobates OR "Costa Rica"	Recovers all records that contain the word "Lithobates" or "Costa Rica": 402 records inasmuch as this will include all records for the entire amphibian fauna of Costa Rica
Combining Boolean operators within the Basic Search box (AND, OR, NOT, quotation marks)	Lithobates AND "Costa Rica" NOT forreri	Recovers only records that contain the word Lithobates AND "Costa Rica", but excludes any record that contains the word "forreri" in any part of the record: 9 records
Printable HELP file

Hyla simplex Boettger, 1901

Class: Amphibia > Order: Anura > Family: Hylidae > Subfamily: Hylinae > Genus: Hyla > Species: Hyla simplex

Hyla chinensis var. simplex Boettger, 1901, Ber. Senckenb. Naturforsch. Ges., 1901 (Wiss. Abhandl.): 53. Holotype: SMF 2626, according to Mertens, 1967, Senckenb. Biol., 48(A): 41. Type locality: "Phuc-Son in Annam", Vietnam.

Hyla simplex — Boulenger, 1903, Ann. Mag. Nat. Hist., Ser. 7, 12: 186; Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 477-480; Tian, Jiang, Wu, Hu, Zhao, and Huang, 1986, Handb. Chinese Amph. Rept.: 58.

Hyla chinensis simplex — Nieden, 1923, Das Tierreich, 46: 201; Bourret, 1937, Annexe Bull. Gen. Instr. Publique, Hanoi, 1937: 21.

Hyla simplex simplex — Fei and Ye, 2000, Cultum Herpetol. Sinica, 8: 71, by implication.

Hyla simplex hainanensis Fei and Ye, 2000, Cultum Herpetol. Sinica, 8: 71, 73. Holotype: CIB 64III4273, by original designation. Type locality: "Wenchang County (110° 70′ E, 19° 62′ N), Hainan Province, altitude 20 m", China.

Hyla (Hyla) simplex — Fouquette and Dubois, 2014, Checklist N.A. Amph. Rept.: 331, by implication.

English Names

Annam Treefrog (Frank and Ramus, 1995, Compl. Guide Scient. Common Names Amph. Rept. World: 58).

South China Tree Toad (Fei, 1999, Atlas Amph. China: 142).

South China Tree Toad (Hyla simplex simplex: Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 256).

Hainan Tree Toad (Hyla simplex hainanensis: Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 256

Distribution

Vietnam (Annam and Tonkin), northeastern Laos, and southern China (eastern Jiangxi, Zhejiang, Guangxi, western Guangdong, and Hainan), 50 to 1500 m elevation. See comment.

Comment

In the Hyla arborea group of Faivovich, Haddad, Garcia, Frost, Campbell, and Wheeler, 2005, Bull. Am. Mus. Nat. Hist., 294: 102 (see comment under Hyla), although it is likely to be a member of the Hyla eximia group (DRF). See Pope, 1931, Bull. Am. Mus. Nat. Hist., 61: 477–480, Liu and Hu, 1961, Tailless Amph. China: 128–130, and (as Hyla chinensis simplex) Bourret, 1942, Batr. Indochine: 224–226. See account by Ye, Fei, and Hu, 1993, Rare and Economic Amph. China: 205. Fei, 1999, Atlas Amph. China: 142–144, provided a brief account, map, and figure. Huang, 1990, Fauna Zhejiang, Amph. Rept.: 53–54, provided an account for Zhejiang populations. Zhang and Wen, 2000, Amph. Guangxi: 66, provided an account for Guangxi. Orlov, Murphy, Ananjeva, Ryabov, and Ho, 2002, Russ. J. Herpetol., 9: 83, provided the Laos component of the range statement. Zhang, 2002, Sichuan J. Zool., 21: 198–199, provided a key to differentiate this species from others in China. Nguyen, Ho, and Nguyen, 2005, Checklist Amph. Rept. Vietnam: 18, provided specific localities for Vietnam. In the Hyla immaculata group of Fei, Ye, Huang, Jiang, and Xie, 2005, in Fei et al. (eds.), Illust. Key Chinese Amph.: 98, who provided a key to the subspecies. Fei, Hu, Ye, and Huang, 2009, Fauna Sinica, Amph. 2: 605–613, provided accounts by species and subspecies, illustrations of holotypes, and spot maps. Fei, Ye, and Jiang, 2010, Colored Atlas of Chinese Amph.: 256, provided brief accounts as Hyla simplex simplex and Hyla simplex hainanensis. Shi, 2011, Amph. Rept. Fauna Hainan: 45–46, provided an account for Hainan. Fei, Ye, and Jiang, 2012, Colored Atlas Chinese Amph. Distr.: 288–289, provided accounts (as Hyla simplex simplex and Hyla simplex hainanensis), illustrations, and a range map. Fei and Ye, 2016, Amph. China, 1: 854–859, provided accounts by subspecies, photographs, and spot maps for China. Li, 2011, Amph. Rept. Guangdong: 37, provided a brief account for Guangdong, China, and photograph. Poyarkov, Nguyen, Popov, Geissler, Pawangkhanant, Neang, Suwannapoom, and Orlov, 2021, Russ. J. Herpetol., 28 (3A): 28, suggested without providing evidence that records from China may actually correspond to Hyla zhaopingensis and that the Laos record requires confirmation.

External links:

Please note: these links will take you to external websites not affiliated with the American Museum of Natural History. We are not responsible for their content.

For additional sources of information from other sites search Google
For images search CalPhoto Images and Google Images
To search the NIH genetic sequence database, see GenBank
For additional information see AmphibiaWeb report
For information on conservation status and distribution see the IUCN Redlist
For related information on conservation and images as well as observation see iNaturalist
For additional information specific to China see Amphibia China
For access to available specimen data for this species, from over 350 scientific collections, go to Vertnet.