Pelobatrachus Beddard, 1908 "1907"

Class: Amphibia > Order: Anura > Family: Megophryidae > Subfamily: Megophryinae > Genus: Pelobatrachus
7 species

Ceratophryne Schlegel, 1858, Handl. Dierkd., 2: 56. Type species: Ceratophryne nasuta Schlegel, 1858, by subsequent designation of Gorham, 1966, Das Tierreich, 85: 15. See discussion by Dubois, 1980, Bull. Mens. Soc. Linn. Lyon, 49: 472. Synonymy (confusing, see comment) with the junior Pelobatrachus by Zeng, Wang, Lyu, and Wang, 2021, Zootaxa, 5004: 39. 

Pelobatrachus Beddard, 1908 "1907", Proc. Zool. Soc. London, 1907: 909. Type species: Ceratophryne nasuta Schlegel, 1858, by monotypy.

Borneophrys Delorme, Dubois, Grosjean, and Ohler, 2006, Alytes, 24: 15. Type species: Megophrys edwardinae Inger, 1989, by original designation. Synonymy with Megophrys by Chen, Zhou, Poyarkov, Stuart, Brown, Lathrop, Wang, Yuan, Jiang, Hou, Chen, Suwannapoom, Nguyen, Duong, Papenfuss, Murphy, Zhang, and Che, 2017, Mol. Phylogenet. Evol., 106: 41. Synonymy with Pelobatrachus by Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 39. 

Common Names

Clay Horned Toad (Qi, Lyu, Wang, Mo, Zeng, Zeng, Dai, Li, Grismer, and Wang, 2021, Zootaxa, 5072: 430). 

Kinabalu Horned Frog (Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 175).

Distribution

Peninsular Thailand and West Malaysia; Borneo (East Malaysia, Indonesia, Brunei), Sumatra, Singapore, Philippines. 

Comment

Resurrected from the synonymy of Megophrys by Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 39, where it had been placed by Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 39. Haas, Das, Hertwig, Bublies, and Schulz-Schaeffer, 2022, Guide to the Tadpoles of Borneo: 175–178, summarized the knowledge of habitat, reproduction, larval morphology and coloration. 

Comment from DRF: The status of Ceratophryne Schlegel, 1858, is confusing, but apparently threatens seniority of the name Pelobatrachus.  Dubois, 1980, Bull. Mens. Soc. Linn. Lyon, 49: 472, regarded it as being a megophryid, due to the subsequent designation of Ceratophryne nasuta Schlegel, 1858, by  Gorham, 1966, Das Tierreich, 85: 15. Dubois, Ohler, and Pyron, 2021, Megataxa, 5: 475), however, apparently regarded Ceratophryne Schlegel, 1858, as an incorrect subsequent spelling of Ceratophrys Neuwied, 1824, without discussion, although it is possible that the information adjacently on the same page associated with Ceratophryne Günther, 1859, was inadvertently transposed during the publication process. Lyu, Zeng, Wang, Liu, Huang, Li, and Wang, 2021, Zootaxa, 4927: 39, considered Ceratophryne Schlegel, 1858, to be a synonym of Pelobatrachus Beddard, 1908, but, without discussion, employed the junior name, Pelobatrachus, as the appropriate name of the genus. The status of the name requires resolution.  

Contained taxa (7 sp.):

Pelobatrachus baluensis (Boulenger, 1899)
Pelobatrachus edwardinae (Inger, 1989)
Pelobatrachus kalimantanensis (Munir, Hamidy, Matsui, Iskandar, Sidik, and Shimada, 2019)
Pelobatrachus kobayashii (Malkmus and Matsui, 1997)
Pelobatrachus ligayae (Taylor, 1920)
Pelobatrachus nasutus (Schlegel, 1858)
Pelobatrachus stejnegeri (Taylor, 1920)

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